172 Mr. J. Miers on several genera hitherto placed in Solanacez. 
of that of Heteranthia: its fruit is capsular as in the Salpiglos- 
sidee, and its seeds contain a terete embryo, curved in an almost 
spiral form. Its leaves are always alternate and deeply pinnati- 
sected, showing an approach to Salpiglossis and Pteroglossis. 
The abortion of three of its stamens is an irregularity of which 
we find a parallel case in Janthe, which only differs m that re- 
spect from Verbascum ; and the deeply laciniated divisions of its 
corolla is another abnormal feature, but this may be considered 
only as a separation of the lobes of the corolla at each sinus, or 
a return to its five normal divisions, with a still farther cleavage 
of each lobe, by an extension in an excessive degree of the inci- 
sions commenced in the emarginatures of all the lobes of the 
border in Salpiglossis, which thus shows a tendency towards the 
laciniated form of the corolla of Schizanthus. 
5. Salpiglossidee.—lI have ventured to remove this tribe wholly 
from the Scrophulariacee for the reasons'that will be here fully 
explained, and as these are founded upon facts in great measure 
new, I may confidently expect that such an arrangement will 
meet with the concurrence of the author of the able monograph 
of this last-mentioned family, who in detailing the characters of 
the tribe in question, as given in the Prodr. DeCand. x. p. 190, 
goes the length of saying, “ subordo Solanaceis capsularibus arcte 
affinis, et forte melius eis adsociandus.” I propose however to 
remove from it several of the genera there associated. They form 
an extremely natural group, distinguished by the very peculiar 
eestivation of their corolla, their didynamous stamens, or where a 
fifth occurs it is invariably sterile, and they are especially eonspi- 
cuous for the remarkable dilatation of the stigma, which at once 
signalizes them from the others. Their place is manifestly 
among the Atropacee, with which they agree in having the ori- 
gin of the pedicels always somewhat lateral in regard to the floral 
leaflet or bract, not decidedly axillary, as in the Scrophulariacee. 
They are all herbaceous plants, generally clothed with viscid 
glandular pubescence, and the campanular portion of the tube of 
the corolla is plicated in estivation ; but the lobes of its border 
are first conduplicate, with the margins always free from those 
of the contiguous lobes, and twisted inwards in a peculiar man- 
ner, for which I have proposed the term reciprocative*, a con- 
dition intermediate between the induplicato-valvate estivation 
of the Solanacee and the imbricate preefloration of the Serophu- 
lariacee ; in order to render this more evident, the accompanying 
* It may be thus defined: stivatio reciprocativa, 7. e. lobi superioris 
exterioris marginibus utrinque induplicatis, loborum alterorum simpliciter 
conduplicatis, 2 sinistralibus dextrorsim, 2 dextralibus sinistrorsim torsive 
convolutis, marginibus Sese applicitis et a contiguis liberis postice spectan- 
tibus, plicaturis antice inclinantibus. 
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