200 Messrs. Hancock and Embleton on the Anatomy of Kolis. 
ramifications as they are termed of the gastric cavity, which are — 
prolonged into the papille. It cannot however be contended 
that the chyle is transuded through the granular or glandular 
part, such as occurs in many of the Kolidide in the papille, 
since it is manifestly a secreting and not an absorbing surface, 
and the current must set from without inwards. Now in #. de-- 
specta the central duct or stem, and its accessory ducts, as well 
as their terminations in the papille, are granular throughout ; 
therefore the fact of the whole apparatus being one for secretion 
precludes the idea that the products of digestion ean pass into 
the system from this organ. This arrangement we see in a still 
more striking manner im several others of the Eolidide, as in 
Hermea dendritica, in which all parts of the much-branched 
hepatic organ are alike granular. In Eumenis marmorata, in 
which they are even follicular throughout, and in Dendronotus 
arborescens*, the central duct is crowded with compound follicles, 
and all the branches are more or less follicular for a short di- 
stance, and then become simply granular; indeed in this genus 
the posterior part of the stomach and the intestine are the only 
parts which are free from the above granular character. We are 
therefore led to conclude that it is from the pyloric end of the — 
stomach and from the intestine that exudation or absorption of 
the chyle takes place, and this conclusion is strengthened by the 
fact, that it is in the intestine that the contents first assume their 
fecal character. We may add also that in Doto, the intestine, 
which is short and wide, is in the interior longitudinally plicated, 
as if thus to increase the extent of the absorbing surface. 
In conclusion then we hope to have shown, that not in any 
of the systems of organs is Kolis notably below the Nudi- 
branchiate type; and we trust that this memoir, if it serve no 
other purpose, will at least assist in rescuing this genus, and 
* In this genus we see an intermediate link between those members of 
the Nudibranchiata which are provided with a concentrated internal hepatic 
organ and the Lolidide, a fact which we pointed out two years ago. The 
central duct is in fact nothing else than a true liver reduced somewhat in 
bulk, but being diffused by its prolongations into the branchial papilla. 
Another intermediate form and still more interesting link between the two 
extremes, as it exhibits the first step in the deviation of the liver from the 
typical state, is seen in Scy/le@a, and which we noticed in a paper communi- 
cated to the Oxford meeting of the British Association. ‘The liverin Seylle@a 
is broken up into several globular masses of convoluted tubes sending off 
minute branches that ramify in the skin and penetrate the branchial tufts. 
In a paper by M. E. Blanchard in the * Annales des Sciences Naturelles’ for 
March 1848, we observe that that gentleman has discovered in Tethys a 
similar arrangement of parts, and points this out as an excellent intermediate 
illustration of the affinities that exist among the different members of the © 
Nudibranchiate group, and we are happy thus to find in his researches a 
corroboration of the fact which we had previously cited for the same end. 
ys 1 
