SCLEROTIA OF UNKNOWN AFFINITY. 267 



neck of living Rliinanthus minor: these bodies begin their 



development in the cambium and bark, which they kill ; after- 

 wards the wood itself mav be attacked. 



Sclerotinia with Botrytis-conidia.^ 



Scl. Fuckeliana ]Je Bary. This Sdrrot'mia is distinguished 

 from all preceding ones by its passing through a Botrytis- 

 conidia stage {Botrytis cinerca). If conidia are sown out on 

 plum-juice gelatine, there appear within fourteen to twenty-one 

 days round groups of sclerotia, which soon 

 give rise to conidia. From such artificially- 

 reared sclerotia I have never succeeded in 

 getting the Peziza-ixnit, so easily cultivated 

 from sclerotia gathered in the open-air {e.y. 

 from vine leaves)." Thus the actual proof 

 that Scl. Fucldiana and Botrytis cinerca are /.vJ;tc"i«na°' i^ptrorof 

 stages in the life of the same fungus is not scieroth?^' (v.^'^'Tubeuf 

 reached by this experiment.^ The two forms ^^^'^ 

 are, however, very frequently met together. 



The sclerotia of Scl. Fuckeliana are produced in the mesophyll 

 of the leaves, also in the parenchyma and epidermis of the 

 host-plants, but never in the wood. Peziza-ixmts, with flat 

 apothecia are produced from them. Sclerotia are found in vine 

 leaves and over-ripe grapes (Fig. 140), especially of the Eiesling, 

 Orleans, and Sylvaner varieties.'* Other plants and fruits may 

 also be attacked. Diseased parts become brown from the 

 effects of the parasitic mycelium, and die off The mycelium 

 can only live parasitic after it has been strengthened by a pre- 

 vious saprophytic existence. Ascospores are thus unable to 

 effect direct infection. The Botrytis-co\\\d(\di seem, however, 

 capable of directly infecting a host-plant, at least I have always 

 succeeded in infecting Conifers successfully with the conidial 

 form Botrytis Dovglasii. 



'See also Botrytis amongst the "Fungi iniperfecti." 



-Brefeld, Heft iv., p. 129, and x., p. 315; Tubeuf, Beitrdge z. Kenntniss d. 

 Baumkrankheiten, 1888. 



^Zopf. (Die Pilze, p. 742) states that Peziza-ivmts may be reared from these 

 sclerotia after they have rested a year. 



^ Muller-Thurgau, "Die Edelfiiule d. Trauben." Landicirlh. Jahrhuch, 1888 

 (Ref. in Botan. Centralhlatt, xxxv., 1888, p. 94). 



