External Characters of Ruminant Artiodnctyla. 223 



compared with its height, the height in the middle line 

 being slightly less than that of the upper lip and much less 

 than the width of the internarial septum. 



As in other members of this subfamily the inguinal 

 glands are absent, there are two pairs of mammce, and the 

 pedal glands are present and constructed as in Oryx. 



The penis (fig. 4, C, D) is remarkable for the thickness 

 and length of the tubular prolongation of the urethral canal, 

 whicli projects some distance, beyond the ovate termination 

 of the glans and is nowhere adherent to it. It rises from 

 the underside of the cylindrical portion of the penis, and, 

 although normally closely applied to the left side of tlie well- 

 defined terminal portion, is in reality separable from it. It 

 is thick at the base and gradually attenuated apically. 



The penis closely resembles that of Addax as described 

 and figured by Garrod (P. Z. S. 1877, p. 10, fig. 18) ; but 

 Gerhard t^s figure of the penis in Addax has a much shorter 

 urethral prolongation, not overlapping the tip of the glans 

 (Verb. Deutsch. Zool. Ges. xvi. p. 153, 1906). 



Genus Hippotragus, Sund. 

 Hippotragiis niger, Ham. (p. 909). 



Specimens examined since 1910 confirm in every parti- 

 cular the facts stated in that year as to the structure and 

 incidence of the cutaneous glands. The new facts here 

 recorded relate to the rhinarium and the penis. 



The muzzle (fig. 5, A, B) is less depressed than that of Oryx, 

 and the sculptured rhinarium is relatively larger and more 

 normal, with a better defined naked tract below the nostrils, 

 a well-developed philtrum mesially grooved, broad at the 

 base, and narrowed below where it reaches the inferior edge 

 of the upper lip ; the nostrils are more expanded with the 

 upper lid more swollen, so that from the anterior asiject the 

 upper edge of the rhinarium appears to be biconvex with an 

 angular median depression and from the dorsal aspect the 

 anterior edge is seen to be transversely truncated. The hairs 

 of the upper side of the nose extend some distance between 

 the nostrils, although not so far as in Oryx, so that the 

 posterior border of the rhinarium is strongly concave. 



T\\e, penis (fig, 4, E) resembles that of j^goryx, described 

 above, in the thickness, length, and freedom of the tubular 

 prolongation of the urethral canal, but the termination of 

 tlie glans is less markedly bulbous. 



The Oryginre, apart from Addax which requires examina- 

 tion, are a remarkably uniform group with respect to the 



