316 The Classification &c. of F. uncia and its Allies. 
It may be added that Matschie’s classification of these 
large Felide as Uncia for leo, tigris, and concolor and Leo- 
pardus for pardus, uncia, and onca (SB. Ges. Nat. Fr. 
Berlin, 1895, pp. 198-199)—a classification which was 
adopted by Trouessart in 1904 (Cat. Mamm., Suppl. p. 265) 
—is quite indefensible both from the zoological and nominal 
standpoints. The valuelessness of Matschie’s opinion on the 
question of the affinities of the species concerned is attested 
by his regarding uncia as a subspecies of pardus and by his 
placing concolor with tigris and leo in a genus from which 
pardus and onca are excluded. It cannot be doubted that 
the relationship between onca and pardus is greater than 
that between wncia and pardus, and that tigris is much more 
nearly akin to pardus than it is to concolor. Yet Matschie’s 
classification implies the precise opposite of these conclusions. 
And as regards his choice of names, uncia by tautonymy, 
let alone the selective actions of Severtzow and Gray, is the 
type of Uncia and Leopardus, rejecting Forskal’s work, is 
not admissible either for pardus or onca or wncia, since none 
of these species was included when it was first proposed by 
Gray in 1842 (Ann. & Mag. Nat. Hist. x. p. 260). It was 
applied to griseus, pictus, ellioti, and horsfieldi. Since one 
of these must be the type, I select griseus, which probably 
connotes a subspecies of pardalis. That Gray intended 
pardus to come into Leopardus is shown by his subsequent 
writings ; but there seems to be no defensible pretext for its 
admission, gratifying as it would be to relegate Leopardus to 
the synonymy of Panthera. ‘ 
The principal cranial differences between Uncia and Pan- 
thera may he briefly contrasted as follows :— 
a, Outer chamber of the bulla very large and involving the 
whole of the anterior portion of its cavity, the line of 
the partition remote from the meatus and running from 
the stylomastoid foramen to a point on the inner surface 
close to the basioccipital suture; a narrow valley be- 
tween the bulla and the glenoid joint; basioccipital 
deeply excavated laterally, a deep pit at the anterior end 
Of the CXCAVATION, 2): .é/ji cal u iiisus vine site elon tele Uneia. 
b. Outer chamber of bulla comparatively small, not involving 
the whole of the anterior portion of the cavity; the 
partition-line close to the meatus and ceasing towards 
the anterior edge of the bulla; a wide valley between 
the bulla and the glenoid joint; basioccipital at most 
shallowly excavated laterally .......0ssecsaesceuns Panthera. 
