and the Genealogy of the Arthropoda, 449 
as belonging to an insect which must have been precisely in- 
termediate between the Orthoptera and Neuroptera. By this 
discovery and that of Hugereon, important and hitherto quite 
unsuspected steps have been made towards the establishment | 
of the genealogical relationship of the order of Insects. 
As regards Hugereon, in order to indicate the position which, 
in my opinion, it must occupy in the genealogical tree, I will 
here reproduce the concluding paragraph of my memoir in 
the ‘ Bia intooraphica,’ 
“If we compare the organization of the recognizable parts 
of our fossil with living forms of Insects, we arrive at the 
oo saga result that we have to do with an animal which 
will not enter into any of our orders of Insects hitherto re- 
garded as so firmly established. Not only M..Tischbein, to 
whose kind intervention I am indebted for the intellectual 
possession of the animal, but also Dr. Hagen, of Kénigsberg, 
to whom I sent it for his inspection and opinion, regarded it 
as an Hemipteron, the latter, however, with this limitation :— 
‘ Probably it constitutes a perfectly new form, which, on ac- 
count of the labium, scarcely agrees with the existing Hemi- 
ptera, but can only be referred to them.’ My own opinion was 
originally the same; but I am now decidedly of opinion that , 
I have an insect before me to which our divisions do not 
apply, and which therefore stands outside our system. The 
Wings, especially, prevent my referring it to the Hemiptera. 
No Hemipteron is destitute of the clavus on the anterior wings; 
and in none do the longitudinal veins-show a tendency to attain 
the inner margin, but all are directed towards the apex of the. 
wing. Moreover there are no Hemiptera with antenne re- 
sembling those of Hugereon. The antenne of Hemiptera are 
of several (7. e. 4—5) joints, or, if we count all the small inter- 
mediate joints (e. g. in Letrichodia), of 8-9 joints; but this 
is the highest number. The form, however, of these joints is 
essentially different from that of the antennal joints of Hu- 
gereon. In the Hemiptera they are long, unequal, and here and 
there furnished with dilatations or other alterations of form ; 
in the latter small and all alike. To this we may add the 
formation of the buccal organs. The rostrum of the Bugs 
consists, as is well known, of a nearly closed multiarticulate 
tube, in which the filiform mandibles and maxille are freely 
moveable. ‘The tube consists of the labium amalgamated with 
the labial palpi. In Hugereon we find all these elements pre- 
sent, but very differently developed. The mandibles and max- 
ille are not filiform, nor does the labium form a tube. And. 
yet it is not difficult to regard this structure of the buccal 
organs as a preliminary step towards the’existing Hemipterous. 
