ADAPTATIONS FOR INTERCROSSING. 



223 



so close at hand, little if any of the pollen of that flower can 



reach the stigmas. These actually get fertilized by pollen 



brought by bees, which come loaded with it from 



other flowers and other plants. Symphyandra differs 



from a true Campanula chiefly in the continued 



cohesion of the five anthers into a tube around the 



style. (Fig. 429, 430.) The pollen is discharged on 



and held by the hairy upper portion of the style. 



Soon after, the corolla expands, the lower 



part of the st} 7 le lengthens, and carries 



the pollen-loaded part out of and above 



the anther-tube, as in Fig. 430 ; lastly, the 



three connivent tips of the style diverge 



and expose the stigmas to pollen mainly 



brought by bees from other flowers. By a 



slight further modification in Lobelia and 



in Compositae, pollen is pushed out of the 



anther-tube by the tip of the style as it 



lengthens, or by the very back of the two 



stigmas, the faces of which, afterwards 



exposed, are not to receive this, but other 



pollen, though it may at times receive some of its own. The 



arrangement in Compositae is here illustrated from Leptosyne 



maritima (Fig. 431-435), a showy plant of Southern California, 



now not very rare in cultivation. The large flowers around the 



FIG. 429. Stamens and pistil of a young, and 430, same from an old flower at 

 Symphyandra pendula 



FIG. 431. Head of flowers of Leptosyne maritima, of the natural size. 



