FRINGE-PINNED GROUP. 517 



istic of the Secondary period, their remains being especially common in the British 

 Lias. In both of the two principal families the tail is of the heterocercal type. 

 In one family, as typified by the genus Palceoniscus, the body is elongated fusiform, 

 and the teeth are slender and conical or straight. On the other hand, Platysomus 

 represents a second family (Platysomatidce), in which the body is rhomboidal, 

 and the teeth in the upper jaw mainly confined to the pterygoid bones obtuse. 

 In both groups the scales are of the ganoid type. 



THE FRINGE-FINNED GANOIDS, Order CROSSOPTERYGII. 



The whole of the members of the subclass under consideration described in 

 the foregoing pages constitute one great order (Actinopterygii), characterised, as 

 mentioned on p. 334, by the fan-like structure of the paired fins, and frequently 

 also of the caudal fin ; the scales being generally of the cycloid or ctenoid type. 

 These fishes form, indeed, the dominant group at the present day ; whereas the group 

 now to be considered is represented only by two existing species referable to as 

 many genera, and is mainly characteristic of the earlier epochs of the earth's 



THE BICHIR. 



history, being abundant even in the Devonian and Carboniferous epochs, since 

 which time it has been steadily decreasing in numbers. These fringe-finned 

 ganoids, as they may be called, have the paired fins lobate, with an internal 

 longitudinal axis belonging to the true skeleton more or less fringed with dermal 

 rays, the caudal fin being either of the diphycercal or heterocercal type. A pair 

 of large jugular plates, bounded in some instances by a series of smaller lateral 

 ones, and an anterior unpaired element, are developed in the branchiostegal 

 membrane to fill up the space between the two branches of the lower jaw, and 

 thus representing the branchiostegal rays of the first order. In all the scales are 

 coated with ganoine, although they may be thin, overlapping, and rounded, or thick 

 and quadrangular. The existing forms have the optic nerves simply crossing one 

 another, a spiral valve in the intestine, and a duct to the air-bladder ; the presence 

 of the latter being also shown in certain extinct types. Next to the sharks and 

 rays, this group is one of the oldest, being well represented in the Devonian. 



The sole existing survivors of this great group of fishes are the 

 Existing 1 Species. 



bichir (Polypterus bichir} of the Nile, and other rivers of Tropical 



Africa, and the reed-fish (Calamoichihys calabaricus) from Old Calabar; these 

 constituting the family Polypteridce, which has no fossil representatives, and 

 probably forms a subordinal group by itself. In this family the notochord is more 

 or less constricted and replaced by ossified vertebrae; the baseosts, or superior 

 supporting elements, are rudimentary, or wanting, in the median fins ; whereas the 



