I. SPECIES INPOraftnON 



1. Classification and ncmenciLature. 



A. S^pecies. 



1. Scientific name. 



a. Bincmial: Hcwellia aquatilis A. Gray. 



b. Full bibliogra^jhic citation: Gray, A. 1879. Prxx:. Am. 

 Acad. 15: 43-44. 



c. Type specimen: Oregon, Multnomah County, Sauvies 

 Island, 1879, Thomas & Joseph Hc^ell 137 . GH. 



2. Pertinent synonyns: None. 



3. Ocninon name: Water HcKx^llia. 



4. Taxan codes: PDCAMOAOIO (The Nature Conservancy) ; 4886, 

 HOWAQU (U.S. Forest Service, Region 1); 17T.L12 (Washington 

 Natural Heritage Program; California Nongame-Heritage 

 Program); HQAQ (Garrison et al. 1976; Soil Conservation 

 Service 1982) . 



5. Size of genus: Monotypic genus. 



B. Family classificaticn. 



1. Family name: Cairpanulaceae. 



2. (Pertinent family synonyms: None. 



3. CCnmDn names for family: Haunetoell Family, Bellflcwer 

 Fcunily. 



C. Major plant group: Dicotyledoneae. 



D. History of knowledge of taxcn: Hcwellia aquatilis was first 

 collected in May, 1879, by two early Oregon botanists, Thonas and 

 Joseph Hcwell. The initial discovery was made in a slough on 

 Sauvies Island, alcxig the Columbia River necur Portland. The 

 initicdly collected material included only submergent, 

 cleistogamous flowers. They returned to a nearby eur^a in August 

 of that year, and collected material becuring well-developed, 

 emergent, chcismogcunous flowers. The specimens were determined to 

 represent a new genus and species by Asa Gray, and it was 

 described in the same yeaur (Gray 1879) . 



Subsequent collections were made in Mendocino County, California 

 in 1928 (Smith and Berg 1988); Clackamas (1892), Marion (1926, 

 1928), and Multnomah (1879, 1881, 1885, 1886) counties, Oregon 

 (OregOTi Natural Heritage Data Base); Clark (1980), Mason (1937), 



