THE PRINCIPLE OF COHESION. 7o 



then becomes a cup, wliicli finally contracts above to 

 form the stjle, just as in Primuiacea\ It is, therefore, 

 unilocular, while a circle of ovules appears on a thick ring 

 of tissue within the base of the ovary. Other circles of 

 ovules appear concentrically and centrifugally. It might be 

 questioned, therefore, whether the ring which carries them 

 were axial or not. I think, however, the same interpreta- 

 tion will apply here as elsewhere ; that is to say, the ovules 

 arise from the place where the bases of the carpels luould 

 have ajjpeared had they been differentiated out of the axis. 



Iq the allied genus Drosera the placentas are strictly 

 parietal, and the ovules, commencing to emerge lialf-way up 

 the wall, appear successively, l>oth upwards and dcwnwards. 

 Now, as they are centrifugal in Diona'a (corresponding to 

 the upward development in Drosera), it looks as if only a 

 portion of the upper half of the carpels were really repre- 

 sented at all. 



In this genus there is a barren central space within the 

 ring of ovules, 2oerhaps representing the termination of the axis. 



That the basal portion only of syncarpous pistils should 

 bear ovules is common enough, and the pla- 

 centas often swell out there to form bosses 

 which we may reasonably conceive as coa- 

 lescing to form the continuous ring character- 

 istic of Dionam. Thus.4cer illustrates how each 

 of the two carpels gives rise to two globular 

 protuberances on which the ovules are borne 

 (Fig. 17). Anemiopsis, as figured by Payer, 

 has a confluent protuberance bearing several 

 basifuofal ovules. Similar multiovular bosses i j.t. i7— Curpois -if 



° o 7 1 c 7 7- • • ^cer (altor Payer). 



occur m boianece and hcropliularinca\ giving 

 the characteristic dumb-bell shape in a transverse section. 

 Now, if we imagine these SAvolleu ovuliferous placentas 



