Rutaceae. 



IV. Apocarpae. 



Capsules apocarpous, carpels discreet. 



Leaves opposite, cobwebby, capsules glabrous. 



Leaves oblong, cobwebby underneath, flowers up to 200 



P. multiflora 

 Leaves ovate to ovate-oblong, cordate, flowers up to 40 



P. Knudsenii 

 Leaves elliptico oblong, curved, concave, chartaceous, flow r ers 



3 to 5 P. barbigera 



Leaves opposite, capsules pubescent. 



Leaves thin chartaceous, obtuse, pale pubescent, capsule 



puberulous P. elliptica 



Leaves ovate oblong, subcoriaceous, tomentulose, capsule with 

 fulvous tomentum P. cinerea 



Pelea clusiaefolia Gray. 



Alani. 

 (Plate 84.) 



PELEA CLUSIAEFOLIA Gray, Bot. U. S. E. E. (1854) 340, pi. 35; Mann. Proc. Bost. 

 Soc. Nat. Hist. X. (1866) 312. et Proc. Am. Acad. VII. (1867) 158, et Proc. 

 Essex Inst. V. (1867) 165; Wawra in Flora (1873) 107; Hbd. Fl. Haw. Isl. 

 (1888) 62; Heller Plants Haw. Isl. (1897) 838. Clusia sessilis Hook, et Arn. 

 Bot. Beech. (1832) 80 (not Forster). Evodia clusiaefolia Drake Del. Cast. 111. Fl. 

 Ins. Mar. Pacif. VI. (1890) 131. 



A small glabrous tree; leaves in whorls of 4 or 3, occasionally 2, obovate or obovate- 

 oblong, rounded or emarginate, with contracted base, thick coriaceous, with a prominent 

 midrib and continuous intramarginal nerve which is close to the edge, shining above, dull 

 underneath, 5 to 12 cm long, 3 to 6 cm wide, on either short petioles of 1 cm or even 

 subsessile, or on petioles of 2.5 cm; flowers in axillary clusters, often cauline, the thick 

 peduncle scarcely 2 mm in length, the pedicels 2 to 4 mm, minutely bracteate at the base; 

 sterile flowers of the same size as the fertile, in the former some of the stamens are as 

 long as the petals and even longer, protruding, the sepals and petals acute, the latter 

 twice as long as the former, ovary glabrous, rudimentary, composed of 4 globose carpels, 

 with apparently no style in the writer's specimens, but small sessile stigma; fertile 

 flowers 4 to 6 mm, the petals more than twice the length of the sepals, stamens rudi- 

 mentary, little longer than the rather depressed ovary, anthers sagittate on broad fila- 

 ments, style 2 mm, with a 4 lobed stigma, the lobes rather thick and blunt; capsule 4 

 lobed, the carpels united to the middle, obtuse or obovate, prominently marked with con- 

 centric wrinkles, one to two seeded, 16 mm in diameter. 



Wawra says of this species that the flowers are hermaphrodite ; this is, however, 

 not the case. All Hawaiian Peleae have fertile and sterile flowers with either one 

 or the other organ rudimentary, making them appear to be hermaphrodite. The 

 male flowers of this species were not known to Asa Gray. 



Wawra in Flora records three forms: fm. a (normalis) from the Waianae 

 Mts., fm. (3 (macrocarpa) and fm. y microcarpa. Asa Gray enumerates two va- 

 rieties ft and y, so does Hillebrand. 



The writer has large material of this species from many localities. It is one 

 of the most common Pelea on Oahu, as well as on other islands, especially on 

 Hawaii in the forests of Puna, near the Volcano Kilauea. 



It is a medium-sized tree reaching a height of 25 to 30 feet in certain localities. 



Specimens from Konahuanui, Oahu, have ovate acute leaves, but also varying 

 tremendously, while others from the Waikane Mts., on the windward side of 

 Oahu, have obovate subsessile leaves. From the same locality the writer col- 



215 



