98 ANTHOCEROS. 



like foot become adapted to the ground, and the chloro- 

 phyllose tissue increased. This course of development 

 for the foot, however, was never worked out, or at least 

 has not survived. This is the highest type of sporophyte 

 found in the liverworts, and gives a hint of the inde- 

 pendent sporophyte of the Pteridophytes soon to be con- 

 sidered. 



The asexual spores of Anlhoceros are formed in the 

 extended portion or capsule, the oldest being at the top 

 and the youngest below Only a cylinder of tissue is 

 used to produce spores, all the remaining part being 

 differentiated to serve other functions. It is evident 

 that we have had a relative, though probably not an 

 absolute, diminution of the amount of sporogenous tissue, 

 and corresponding increase of sterile tissues from the 

 lowest to the highest liverworts. This has been accom- 

 panied by increase in the complexity of the sporophyte 

 body, and constant approach toward independence. 1 



1 See article by Bradley Moore Davis on "The Origin of the Sporo- 

 phyte," already cited in connection with CoUochate. 



