SINCLAIR: MARSUPIALIA OF THE SANTA CRUZ BEDS. 371 



form, which does not extend to the distal margin of the bone, as in the 

 latter genus. The facet is oval in outline and slightly concave ; that for 

 the navicular, which immediately adjoins it, is triangular and convex. 

 The cuneiforms resemble those of Sarcophilus rather than Thylacynus. 

 The inner cuneiform is laterally flattened, supporting distally an oval, con- 

 cave facet for the metatarsal of the hallux. Laterally it is in contact with 

 the mesocuneiform by a narrow elliptical facet. The mesocuneiform is a 

 small, oblong element, much shorter than the adjacent bones, so that the 

 proximal end of the second metatarsal is received between them, articu- 

 lating with the mesocuneiform by a concave oval facet, and with the 

 ectocuneiform by a small plane facet confined to the dorsal margin of that 

 bone. The outer cuneiform differs from that of Thylacynus and resem- 

 bles Sarcophihis in lacking the large plantar process present in the former 

 genus. It is in contact with the cuboid by a large triangular facet and 

 has a very small facet for the fourth metatarsal. The facet for the meso- 

 cuneiform is narrow and sigmoid in dorso-plantar section. 



The metatarsals are shorter and proportionately heavier than in Thyla- 

 cynus. The hallux is vestigial, its distal end terminating in a rugose 

 knob, which did not support phalanges. Proximally, the shaft bears a 

 broad, plane facet for contact with metatarsal II. The third metatarsal 

 is little more than half as long as the corresponding element in Thyla- 

 cytms. The proximal end lacks the prominent plantar tubercle present in 

 the latter and the articular surface for the ectocuneiform is more convex. 

 The shaft is slightly curved and considerably flattened antero-posteriorly. 

 The trochlear surface extends dorsally about as far as in Thylacynus. 

 As in that genus, the keel is confined almost entirely to the plantar sur- 

 face. The fourth metatarsal of the right pes is preserved. It is consid- 

 erably longer than the third metatarsal. The proximal end is wider and 

 more strongly convex in dorso-plantar section than in Thylacynus. 



The antero-posterior shortening and especially the transverse flattening 

 of the trochlear surface of the astragalus are believed to indicate a planti- 

 grade gait. 



Restoration (PI. LXI, fig. 2). The restored skeleton shows effec- 

 tively the large head, long neck and short limbs characteristic of all the 

 Santa Cruz marsupial carnivores. The lack of proportion between the 

 femur, tibia and hind foot is at once apparent. The length of the tail is 

 conjectural, but the error is probably in underestimating rather than in 



