124 HETEROSTYLED DIMORPHIC PLANTS. CHAP. III. 



fore, is in all respects a well-characterised heterostyled 

 species. 



^GIPHILA ELATA (VERBENACE^). 



Mr. Bentham was so kind as to send me dried flowers 

 of this species and of 2E. mollis, both inhabitants of South 

 America. The two forms differ conspicuously, as the deep- 

 ly bifid stigma of the one and the anthers of the other 

 project far above the mouth of the corolla. In the long- 

 styled form of the present species, the style is twice and a 

 half as long as that of the short-styled. The divergent 

 stigmas of the two forms do not differ much in length, nor 

 as far as I could perceive in their papillae. In the long- 

 styled flowers the filaments adhere to the corolla close up 

 to the anthers, which are enclosed some way down within 

 the tube. In the short-styled flowers the filaments are free 

 above the point where the anthers are seated in the other 

 form, and they project from the corolla to an equal height 

 with that of the stigmas in the long-styled flowers. It is 

 often difficult to measure with accuracy pollen-grains 

 which have long been dried and then soaked in water ; but 

 they here manifestly differed greatly in size. Those from 

 the short-styled flowers were to those from the long-styled 

 in diameter in about the ratio of 100 to 62. The two forms 

 of 2E. mollis present a like difference in the length of 

 their pistils and stamens. 



Flowers of this bush were sent me from St. Catharina 

 in Brazil, by Fritz Miiller, and were named for me at 

 Kew. They appeared at first sight grandly heterostyled, 

 as the stigma of the long-styled form projects far out of 

 the corolla, whilst the anthers are seated halfway down 

 within the tube ; whereas in the short-styled form the an- 

 thers project from the corolla and the stigma is enclosed in 

 the tube at nearly the same level with the anthers of the 

 other form. The pistil of the long-styled is to that of the 

 short-styled as 100 to 60 in length, and the stigmas, taken 

 by themselves, as 100 to 55. Nevertheless this plant cannot 

 be heterostyled. The anthers in the long-styled form are 

 brown, tough, and fleshy, and less than half the length 

 of those in the short-styled form, strictly as 44 to 100; and, 



