CHAP. V. HETEROSTYLED DIMORPHIC PLANTS. 37 



than might have been expected. Secondly, nine capsules 

 from another equal-styled plant, which had not been pro- 

 tected from insects, but probably was self -fertilised, gave 

 an average of 45.2 seeds, with a maximum of 58. Thirdly, 

 another plant which had a very short pistil in 1865, pro- 

 duced spontaneously many capsules, six of which con- 

 tained an average of 33.9 seeds, with a maximum of 38. In 

 1866 this same plant had a pistil of wonderful length ; for it 

 projected quite above the anthers, and the stigma resem- 

 bled that of the long-styled form. In this condition it pro- 

 duced spontaneously a vast number of fine capsules, six of 

 which contained almost exactly the same average number as 

 before, viz. 34.3, with a maximum of 38. Four flowers on 

 this plant, legitimately fertilised with pollen from a short- 

 styled cowslip, yielded capsules with an average of 30.2 

 seeds. Fourthly, another short-styled plant spontaneously 

 produced in 1865 an abundance of capsules, ten of which 

 contained an average of 35.6 seeds, with a maximum of 

 54. In 1866 this same plant had become in all respects long- 

 styled, and ten capsules gave almost exactly the same aver- 

 age as before, viz. 35.1 seeds, with a maximum of 47. Eight 

 flowers on this plant, legitimately fertilised with pollen 

 from a short-styled cowslip, produced six capsules, with the 

 high average of 53 seeds, and the high maximum of 67. 

 Eight flowers were also fertilised with pollen from a long- 

 styled cowslip (this being an illegitimate union), and pro- 

 duced seven capsules containing an average of 24.4 seeds, 

 with a maximum of 32. The fifth and last plant remained 

 in the same condition during both years: it had a pistil 

 rather longer than that of the true short-styled form, with 

 the stigma smooth, as it ought to be in this form, but 

 abnormal in shape, like a much-elongated inverted cone. 

 It produced spontaneously many capsules, five of which, 

 in 1865, gave an average of only 15.6 seeds ; and in 1866 ten 

 capsules still gave an average only a little higher, viz. of 

 22.1, with a maximum of 30. Sixteen flowers were ferti- 

 lised with pollen from a long-styled cowslip, and produced 

 12 capsules, with an average of 24.9 seeds, and a maximum 

 of 42. Eight flowers were fertilised with pollen from a 

 short-styled cowslip, but yielded only two capsules, con- 

 taining 18 and 23 seeds. Hence this plant, in function and 

 partially in structure, was in an almost exactly intenne- 



