154 CRUCIFERyE. Dentaria. 



D. maxima, Nutt. Tnbers near the surface and stems reclined in fruit : leaves two or 

 three, alteruate, 3-foliolate ; leaflets ovate or oblong-ovate, coarsely toothed and somewhat 

 cleft or lobed, 1 or 2 inches long: pods as in i>. laciniata; seeds round-oblong; cotyledons 

 uneciual, the smaller cuneate-obloug, half as wide as the larger ; radicle acute and curved 

 cleft to the middle. — Gen. ii. 66 ; Gray, Gen. 111. i. 138, t. 56. D. laciniata, var. 5, Torr. & 

 Gray, Fl. i. 86. Cardamine maxima, Wood, 1. c. — Vermont, Morgan, to Pennsylvania aud 

 Western New York ; said to bloom two weeks later tiian D. laciniata. Nuttall's original 

 specimens from Pennsylvania and W. New York are described as two feet tall and with five 

 to seven leaves. Nothing corresponding to this appears to have been found since. The 

 single small specimen, so named by Nuttall in the herb. Brit. Mus., from Pennsylvania, 

 has a pair of soj)arafe ternate leaves aud probably belongs to this species or possibly to 

 D. diphijila. 



++ ++ Western species. 



D.* macrocarpa, Ni'tt. Glabrous or slightly pubescent : stems simple, 4 to 15 inches high ; 

 joints of the rootstock about an inch long: leaves 1 to 3, approximate, shortly petiolate, 

 palmately or pinnately 3-5-parted or -divided, the segments linear to oblong, entire, obtuse 

 or acute, ^ to 2 inches long; basal leaves sometimes merely lobed or cleft; the leaflets 

 sessile or petiolulate, often 3-5-lobed or -toothed: raceme usually nearly sessile: flowers 

 purple or rose-color; pods one or two inches long (including the style, usually 3 lines long) 

 and a line l)road ; stigma capitate aud entire ; seeds oblong ; cotyledons somewhat unequal ; 

 the oblique radicle cleft to the base. — Nutt. in Torr. & Gray, Fl. i. 88; Brew. & Wats. 

 Bot. Ciilif. i. 30. D.tenella, Brew. & Wats. 1. c. as to pi. Calif.; Wats. ms. of present work, 

 not Pursh. D. (/emmala, Wats, as to pi. of Howell, Pacif. Coast PI. 1887 (not as to type 

 which was later identified with D. tenella). Cardamine Nuttallii, Greene, Bull. Calif. 

 Acad. Sci. ii. 389. C. ^remHia/n, Greene, Pittonia, i. 162. — N. California (Plumas aud 

 Siskiyou Counties) to Brit. Columbia, Li/all.^ 



Var.* pulcherrima, Robixson, n. var. Flowers larger than in the type ; petals 6 to 

 8 lines long, 4 to 5 lines broad. — Cardamine •pulcherrima, Greene, Erythea, i. 148. — Mosier, 

 Oregon, T. Howell. Very nearly related species, if distinct at all, are the following: 

 CaudAmine sinuIta, Greene, 1. c, with suborbicular sinuate-dentate radical leaves aud roots 

 said to be tuberous, from. Crescent City, Calif., T.IIoicell, and (?) Cow Creek Mts., Oregon, 

 Henderson; also C. quekcetouum, Howell, Erjiihea, iii. 33, witli radical leaves 3-foliolate; 

 leaflets ovate to elliptic-oblong, dentate, from Silverton, Oregon, T. Howell. In their flowers, 

 young fruit (so far as known), cauline foliage, and general habit, these plants sliow such 

 a close resemblance to each other and to more robust forms of D. macrocarpa, that the 

 specific distinctions, derived chiefly from the subdivision of the radical leaves (iu this genus 

 notably inconstant), appear very doubtful. Good specimens of the roots (not at hand) may 

 furnish better distinctions. 



D.* tenella, Pursh. Rootstock bearing small irregular tubers : basal leaves simple, rouud- 

 cordate, coarsely crenate or sinuate, one or two inches broad ; the petiole bearing usually 

 several clusters of bulblets : stem 6 to 12 inches high, with one or two nearly sessile 3-folio- 

 late leaves, sometimes bulbiferous in the axils ; leaflets linear-oblong or linear, obtuse, entire, 

 ^ to 2i inches long: raceme sessile or shortly pedunculate : flowers rose-color: pods an inch 

 long and a line wide, with a slender style tipped with a broad distinctly lobed stigma. — 

 Fl. ii. 439 ; Torr. & Gray, Fl. i. 87. D. 'tenuifolia. Hook. Fl. Bor.-Am. i. 46, not Ledeb. — 

 Banks of the Columbia, Lewis ; Washington, KlikitafCo., Suksdorf, Upper Nesqually Val., 

 Alic.r). 



D. Calif ornica, Nutt. Tubers of the submoniliform rootstock mostly small : stem ^ to 2 

 feet high, rather stout, simple or branched above, glabrous or slightly pubescent : foliage 

 very variable ; basal leaves entire or 3-foliolate ; the leaflets petiolulate, suborbicular, cune- 

 ate to subcordate at l)ase, sinuate or coarsely toothed ; cauline 2 to 4, mostly shortly petio- 

 late and above the middle of the stem, 3- or pinnately 5-foliolate, rarely simple or lobed ; 



1 The treatment of D. macrocarpa and the following species has been revised in the light of more 

 copious material. Nuttall's species was unfortunately characterized as having 3-foliate radical leaves 

 with "reniform " leaflets. It is stated that the species was founded upon a single specimen, and a 

 plant, so labelled by Nuttall himself,, is now in herb. Brit. Mus., and is (ace to Dr. Watson) of the 

 species here described. 



