rpz PHANEROGAMS. 



nucleus, while the subjacent basal part becomes the funiculus. As the integuments 

 arise, the curvature becomes gradually stronger, and the nucleus becomes inverted 

 even bei"ore the outer integument has entirely developed. This latter is there- 

 fore not formed on the side next to the raphe, but clothes all the free part of the 

 ovule, right and left of the raphe (Fig. 366, V, VI, VII). Cramer was the first to 

 point out that anatropous ovules may originate in another way (and this is probably 

 the most common case), the ovule developing as a secondary lateral projection 



Fig. 366.— /—K// stages of development of the ovule of Orchis militaris (x 550) ; I—VI seen from the side in longi- 

 tudinal section, VII from the front, the funiculus being behind, VIII a horizontal section of/ ; xx the axial row of cells, 

 the upper one of which becomes the embryo-sac e, f the funiculus, ii the inner, ai the outer integument, K the nucleus, 

 es the micropyle, h an intercellular space : in VII the embryo-sac e has completely replaced the tissue of the nucleus. 



beneath the apex of the young conical funiculus, and curving backwards subse- 

 quently towards the base of the latter. This inversion takes place while the single 

 or the inner integument is enveloping the nucleus from the summit of the funi- 

 culus ; the second integument, if there be one, then similarly clothes the free part 

 (see Fig. 367, B, C). Kohne^ has indeed thrown some doubt on the actual lateral 

 origin of the nucleus, not only in Compositse, but also in Solanum, Hedera, Fuchsia, 

 Begonia, &c. I have, however, had the opportunity, not only previously, but also 

 more recently, in Grigorieff's researches on Compositae, of observing a number of 



Kohne, Uebcr die Bliithenentvvickelung bei den Compositen. Berlin 1866. 



