EULOTA. 201 



Genitalia : penis extending into an epiphallus which sometimes 

 has, sometimes has not, a flagellum. Dart sack containing a round 

 or flat dart, and either borne on atrium or higher on vagina. Mucus 

 gland inserted on dart sack, or on an empty accessory sack communi- 

 cating with dart sack, and consisting of one or many sacculated or 

 glandular, long or oval branches, bound closely together and to the 

 dart sack. Spermatheca oval or globose, on rather a long duct, 

 which lacks diverticulum. 



Distribution, Central Europe (one species) to Japan, south to E. 

 Indian Archipelago. Especially characteristic of Eastern Asia. 



This genus differs widely from the European dart-bearing Helices 

 in having the one (often many-branched) mucus gland inserted on 

 the dart sack or on an accessory empty sack, and in the structure of 

 the gland itself. It is more closely allied to the American genera 

 Epiphragmophora and Cepolis in the structure of the mucus gland. 



Eulota, as herein understood, comprises a great variety of shell- 

 forms and a large number of species; including, indeed, a consider- 

 able majority of the East Asiatic Helices. As in all other large gen- 

 era of Helices, the shell varies from globular to lens-shaped (see in- 

 troduction to this volume) ; and the several stages of contour, each 

 represented by a numerous progeny of species, have received names 

 which some writers use in a subgeneric, some in a generic, sense. In 

 my opinion, the former is the more philosophic view, as the shell 

 characters fade from one group to another, offering no sharp line of 

 demarkation throughout the genus, so far as I can see. As to the 

 anatomy, my dissections (a part of which are shown on plates 65, 

 66) tell clearly that no grounds for a division of the group into two 

 or more genera can be based thereon, unless the forms in which the 

 penis has a flagellum be separated generically from those lacking 

 this organ ; and I do not think it likely that the examination of 

 more material will add to the value of this feature. At all events, 

 I can find no character of shell or soft parts correlated with it, and 

 we are hardly justified in founding a genus on a single peculiarity, 

 unless it be one of greater value than this. Like Polygyra, Thersites 

 or Helix s. str., the various anatomical divergencies, except as to the 

 flagellum, are fully covered by intermediate forms ; although, as a 

 whole, the genus is characterized by well-marked peculiarities which 

 would enable one to identify any of its members by an inspection of 

 the genital system alone, without the assistance of shell characters. 



The main anatomical divergencies may be tabulated as follows 

 for the species now known anatomically : 



