I 



I 



ORCHIDACEAE (ORCHIS FAMILY) 305 



Oypripedium, of two stamens and the rudiment of a third, variously coakscent 

 with the style. Anther 2-celled, each ceil containing one or more masses oi 

 pollen (pollinia), or the pollen granular (in Cypripedium) . Stigma visciil or 

 (in Cypripedium) rough. Fruit a 1-celled 3-valved capsule. Flowers solitary, 

 racemed, or spiked, often showy, each flower usually subtended by a bract! 

 Leaves parallel-nerved, solitary, or several and alternate, sometimes api)areiitly 

 opposite or whorled. Perennials, often with corms or with tuberoid roots ; 

 sometimes rootless saprophytes. —A cosmopolitan family comprising about 7000 

 species largely dependent on insects for pollination. 



I. Fertile anthers 2. 



Tribe I. CYPRIPEDIeAE. Perfect anthers lateral, the sterile one forming a dilated fleshy append- 

 age above the terminal stigma. Pollen granular, not in masses. 



1. Cypripedium. Stems more or less leafy. Perianth spreading ; lip an inflated sac. 



IL Fertile anther solitary. 



* Anthers persistent. 

 Tribe n. OPHRYDEAE, Pollinia prolonged at the base of the anthers into filaments or caudicles 

 •which are attached to viscid disks or glands. 



2. Orchis. Viscid disks contained in a pouch, or bursicule, of the rostellum. 



8. Habenaria. Viscid disks naked, not contained in a pouch, or bursicule, of the rostellum. 

 * * Anthers caducous or readily detachable. 

 Tribe III. NEOTTIeAE. Pollen-masses usually soft or granulose. 



-J- Anther terminal. 



4. Pogonia. Lip without hypochil, free. Column not winged. 



5. Calopogon. Lip without hypochil, free. Column winged at apex. 



6. Arethusa. Lip without hypochil, united to the base of the gynostemium. Column winged 



to the base. 

 1. Serapias. Lip provided with a hypochil. 



•«- -t- Anther dorsal. 

 ++ Upper sepal and petals connivent or lightly adherent. 



8. Spiranthes. Pollen waxy or powdery, not divided into a large number of definite masses. 



Lip with appendages at the base ; not saccate. 



9. Epipactis. Pollen divided into a large number of definite masses. Lip sa<Jcate, unappendaged. 



++ ++ Sepals and petals free. 



10. Listera. Lip retuse or cleft. 



Tribe IV. EPID^NDREAE. Pollen-masses smooth and waxy. 

 «= Pollen-masses 4, unappendaged. 

 a. Leafless plants. 



11. Corallorrhiza. Plants brownish or yellowish, with coralline rhizome* 



5. Plants with leaves. 



12. Malaxis. Lip not saccate, cordate at base. Leaves several 



13. MicrostyliS. Lip not saccate. Leaf solitary on the stem. 



14. Liparis. Lip not saccate, obovate. Leaves several. 



15. Calypso. Lip saccate. Leafsohtary. 



16. Aplectrum. Lip not saccate. Leaf solitary from a tuber. 



= = Pollen-masses 4, each attached by a very short filament to the viscid disk or gland. 



17. Tipularia. Flowers small, greenish, in a many-flowered raceme. Lip 8-lobed. 



«= «= = Pollen-masses 8. 



18. Hexalectris. Leafless plants. 



gray's manual — 20 



