MUTATION'S, VARIATIONS, AND RELATIONSHIPS (JF THE OENOTHERAS. I I 



OENOTHERA SCIXTILLAXS. 



0. scintiHans was first seen to originate as a mutant in 1888 and has appeared 

 in this manner in 14 individuals at different times in De \'ries's cultures. As 

 previously described in this paper, it has appeared in 4 specimens in the Xew 

 York Botanical Garden during the present year. 



0. scintillans is to be noted as an ever-sporting form which in some strains 

 gives a progeny 60 or 70 per cent of which is composed of lata, oblonga, laniarck- 

 iana, and nanella, while in other strains but ,30 per cent will be included in 

 these forms. The forms thus derived from scintillans are in no wise different 

 in their hereditary qualities from the same types derived from other and pure 

 lineages. Cultures were made from seeds furnished by Professor De \r\ts 

 from a strain which was supposed to yield 60 to 70 per cent of the other types 

 named. 



Seventy-eight individuals were obtained from the culture, 15 of which resem- 

 bled .Tcn?/;7/(7».^-, 46 laniarckiaiia, 16 oblonga, and i was present which did not 

 reach maturity, but suggested some of the incomplete forms mentioned in ' ' Die 

 Mutationstheorie," pvohahly subli)icaris. In the seedlings ,3 months old, the 

 rosettes were 14 to 17 cm. in diameter, dense, and spreading Hat on the ground ; 

 outer leaves, 6 to 7 cm. long, blades about 5 cm. long, 2.-, to 3 cm. wide, 

 oblong-obovate, widest about the middle, obtuse or acutish at the apex, 

 tapering gradually to the white-margined petiole, blue-green, shining, puber- 

 ulent, with occasional small red spots above, glabrous or nearly so, and lighter 

 green beneath, somewhat brittle. At 6 months of age the rosettes were 

 dense, the leaves lanceolate, obscurely and minutely denticulate, green and 

 shining, puberulent on margin and sparingly on upper surface, midvein broad, 

 petioles variously winged. 



The adult plant was short, with few branches which were fairly rigid and 

 ascending. The lower part of the stem appeared to be terete or nearlv so, 

 while the upper portion was angled and channeled. The stem and branches 

 were hirsute with spreading hairs. 



The stem-leaves were finelv pubescent on both surfaces and much crinkled 

 on the margins and midribs; irregularly toothed, oblong, 7 to 9 cm. long, 30 to 

 35 mm. wide, abruptly tapering at base, acutish at the apex. The petioles 

 were short and the entire organ was a deep shining green. 



Conic portion of the bud 15 to 20 mm. long, finely pubescent, with short, 

 spreading hairs. Free tips 5 mm. long, erect. Hypanthium 30 mm. long, and 

 reflexed calyx-lobes 25 cm. long. Ovary 6 to 7 mm. long. 



The petals are thin, 25 mm. long and 30 mm. in width, barely emarginate. 

 Filaments 15 to 18 mm. long, very slender and strongly upcurved. Anthers 

 7 to 8 mm. long. Pistils slightly longer than the stamens ; lobes of the stigma 

 very slender, 4 to 5 mm. long (plate 2). 



