118 C. WINKLEll. THE CENTRAL COUlteE 



lateral fillet (Plate XXII fig. 28 F) of which it forms the greatly 

 atrophied most medial part. 



More distally, where the V th spinal root leaves the cord, its 

 atrophy at the contra- lateral side also seems evident (fig. 28 E). 

 In still more distal sections this is no longer the case and the region 

 between the oliva superior and the fifth spinal root it even is smaller 

 at the operated side (fig. 28 C). 



To interprete the fact I remember that the ascending spino- 

 cerebellar tract has lost fibres. I have shown its atrophy at the same 

 side of the operation , there where it is lying free at the surface of 

 the superior pedunculus cerebelli (fig. 28 F). 



At the opposite side, there is again found atrophy of another 

 part of the lateral fillet and of the ,,aberrirendes Seitenstraugbiindel. 

 This contra-lateral atrophy is found combined with an atrophy of 

 transverse ventral fibres crossing the raphe at the operated side. 

 (Plate XXIII fig. 28 F.) In this distal part of the decussatio ventralis 

 tegmenti (FoiiEL) there is also found degeneration in transverse 

 fibres with MARCHI. (Plate XIX fig. 25 A D). 



It is therefore doubtful whether this ,,aberrirendes Seitenstraug- 

 bundel" that MONAKOW has isolated after transverse section of the 

 oblongata (the ventral ascending spino-cerebellar tract having totally 

 disappeared in such cases) contains only a rubro-spinal descending 

 tract. There must still be found fibres of another origin. 



The current view is that the rubro-spinal tract issued from the 

 nucleus ruber after crossing in the ventral decussatio tegmenti 

 (FoREL) runs downward. Without contact with the red nucleus, 

 there must still exist a crossed portion of that ,,aberrirendes Seiten- 

 strangbiindel" ascending to the medial part of the lateral fillet, and 

 thence in the ventral fibres under the nucleus quadrigeininum 

 posticum passing upward. 



After all, there are entering two important systems into the lateral 

 fillet without reckoning the root-fibres - - that accompany them. 



The one, originating in the contra-lateral tuberculum acusticum, 

 both olivary nuclei, and the same sided nucleus ventralis lemnisci 

 lateralis, are not yet all medullated in the elder foetus or at birth. 

 They go through the fibres of MONAKOAV, the stratum c of the 

 systema ventrale et intermedium into the lateral fillet. 



The other originates in the contralateral nucleus of DEITERS, the 

 contralateral nucleus ventralis N. octavi, the contralateral nucleus 

 griseus nervi descendentis, perhaps in the same-sided nucleus trape- 

 zoides. It passes through the ventral, intermedial and dorsal system 

 of the nervus octavus and is medullated at birth. 



