Nerves, Spinal, Coed, Brain and Other Ganglia 85 



2. The lateral columns, included between the anterior and posterior 

 horn on each side. 



3. The anterior columns, included between the anterior horns, and 

 separated by the anterior fissure. 



In the columns of the cord several distinct groups of fibers are dis- 

 tinguishable. The most important of these are as follows: the numera- 

 tion beginning at the dorsal fissure [Fig. 71]. 



I. Ascending (». e., conducting towards the brain), (a) The greater 

 part of the posterior columns; specifically, the column of Burdach 

 {posterolateral tract or fasciculus cuneatus) , and the column of QoII 

 (postero-mesial tract or fasciculus gracilis). (b) Lissauer's tract, 

 lying at about the apex of the posterior horn, (c) The direct cerebellar 

 tract {dorsal cerebellar tract; fasciculus cerebello-spinalis ; Flechsig's 

 tract), and the anterolateral ascending tract {fasciculus antero-later- 

 alis superficialis ; Goiver's tract), occupying the superficial portion of the 

 lateral column. 



The fibers in the column of Goll and Burdach and Lissauer's tract are 

 axons of the posterior roots, i. e., whose cell-bodies are in the spinal 

 ganglia. The fibers in the direct cerebellar tract have their cell-bodies in 

 Clark's Column; a column of cells in the posterior horn. The location 

 of the cell-bodies of the fibers in Gower's tract have not been definitely 

 identified. 



II. Descending, (a) The septo-marginal tract, lying in the edge 

 of the posterior column close to the dorsal fissure, (b) The comma 

 tract, lying within Burdach's column, close to Goll's. (c) The crossed 

 pyramidal tract {fasciculus cerebro-spinalis lateralis), and the pre- 

 pyramidal tract {rubrospinal tract; von Monakow's tract) lying in the 

 lateral column, (d) The spino-olivary column {bundle of Helwig) 

 lying superficially opposite the anterior horn, (e) The anterolateral 

 descending tract {vestibulospinal tract; marginal bundle of Lowen- 

 thal), marginally in front of the antero-lateral ascending tract, (f) The 

 direct pyramidal tract {fasciculus cerebrospinalis anterior), along the 

 border of the anterior fissure. The fibers in the pyramidal tract are axons 

 of cell-bodies in the cerebral cortex. The cell-bodies of the fibers in the 

 other descending columns lie in the medulla, pons, cerebellum, or mid- 

 brain, or gray columns of the cord. 



III. Spinal interconnecting. The basic bundles, anterior and 

 lateral {fasciculus anterior proprius and fasciculus lateralis proprius), 

 are strands of fibers of cells in the gray matter of the cord, serving to con- 

 nect the different segments of the cord with one another. 



