BRACHIOPODA. 133 



Neither can it lie assigned to Orbiculoidea without opening that genus to the 

 reception of heterogenous forms. For convenience, therefore, it is desirable to 

 indicate the close but subordinate relations of tliis species* to Orbiculoidea, 

 by the use of the new term QEhlertella. 



The muscular impressions in Oroiculoidea are extremely faint and rarely 

 discernible at all. We have seen no specimens showing them as clearly as the 

 pedicle-valve of D. nitida, figured by Mr. Davidson and referred to above, where 

 they are resolvable into anterior and posterior adductors, situated close together 

 near the extremities of the pedicle-area, the posterior pair being at the distal 

 extremities of long progressive scars radiating from near the apex. This 

 arrangement of the scars indicates a general agreement with the muscular 

 anatomy of Discinisca lamellosa. 



Orbiculoidea Randalli, Hall, a remarkably large species, known only from a 

 single rather imperfect interior of the pedicle-valve, from the Hamilton group 

 of New York, shows a slight median septum passing from the apex forward, 

 comparable to that in Discinisca lamellosa ; from this radiate a series of furrows, 

 probably of vascular nature, which lie only within the inner lamellae of the shell, 

 apparently not interrupting the external ornamentation. It will be interesting 

 to learn whether this shell is congeneric with Orbiculoidea Morrisi, Davidson. 



Occasionally, in the brachial valves of American species, are seen two 

 extremely faint ridges beginning near the apex and approaching each other at 

 their anterior termination, nearly meeting the inconspicuous median septum. 

 (See Plate IV f, fig. 22 ; also, Barrande's figure of Discina reversa, Keyserling, 

 Systeme Silurien, vol. v, pi. 95, fig I, 2 a.) In a large form from the Hamil- 

 ton group of New York, usually confounded with 0. grandis, Vanuxem, these 

 characters reach an extraordinary development and become most prominent 

 features of the interior, beginning just in front of the apex, and, as they con- 



* The mode of presei-vation of this species frequently leads to deception in regard to its pedicle-charac- 

 ters. When the two valves are preserved tog-ethnj', the breaking away of the upper i)ortion of the larger 

 valve often leaves its marginal poi-tion surrounding the lower valve and closing the pedicle-aperture. In 

 many such cases it is extremely difficult to distinguish the parts belonging to the resi^ective valves on ac- 

 count of the thinness of the shell, and we have taken pains to accumulate a very large representation of the 

 species in order to fortify our conclusions. For much of the material at our disposal we are indebte<l to the 

 favor of Professor C. L. Hbrbick, of Cincinnati. 



