POLLINATION 61 



carpels dichogamy, and to the movements of the different 

 parts of the flower and the significance of these move- 

 ments in pollination. " Since very many flowers are 

 dioecious/' he says, " and probably at least as many 

 hermaphrodite flowers are dichogamous, Nature appears 

 not to have intended that any flower should be fertilised 

 by its own pollen/' There is the problem of cross pollina- 

 tion stated ; it was left for Charles Darwin to answer the 

 question why this should be so. You will thus see that 

 the labours of Koelreuter and Sprengel threw an entirely 

 new light on the morphology of the flower, and placed in 

 their appropriate niches all the isolated observations on 

 the movements of stamens, of styles, of peduncles, and 

 so forth that had puzzled the earlier naturalists. 



