56 THE NORTH AMERICAN SPECIES OF DROSOPHILA. 



The curious produced eyes of some of the males of Z. dispar caused 

 Wiedemann to describe the species as a diopsine. They do not occur in 

 the female of this species, or in either sex of the other species. Similar 

 eyes are to be found in some undescribed Neotropical species of Chymomyza, 

 close to C. procnemis. The present group is not close to Chymomyza in 

 other respects. 



The two American species have both been bred from fungi. 



Dettopsomyia Lamb. 1914. Trans. Linn. Soc. London, 16, 349. 



Arista plumose; two orbitals, upper reclinate, lower proclinate; postverticals crossed; 

 vibrissa; and ocellars present ; eyes pilose; carina large and broad; two dorsocentrals, plus 

 two small presutural ones; acrostichal hairs in two rows; two humerals; one presutural; 

 two pairs of scutellars, posterior ones crossed; two sternopleurals ; costa reaches apex of 

 fourth vein, twice broken, distal break deep, milichiid-like; costal index a little less than 1.0. 



There is a single described species, D. formosa Lamb, from the Seychelles. 

 Camilla Haliday. 1838. Ann. Nat. Hist., 2, 188. 



Arista plumose; orbitals disposed as usual, middle one minute; postverticals medium 

 size, convergent; vibrissa; present; two notopleurals; mesopleurae bristly; two dorso- 

 centrals; no prescutellars; many acrostichal rows of hairs; presuturals long; two pairs of 

 scutellars; costa twice broken; anal vein absent; anal cell open at apex; auxiliary vein 

 rudimentary; no preapicals on third tibise. 



The type species is the European C. glabra Fallen. Seven species are 

 described, as follows: 



Patearctic: C. glabra Fallen, Europe, Canary Islands; C. acuHpennis Loew, Europe. 

 Etliiopian: C. africana Bezzi, Kongo. 



Oriental: C. coeruleifrons de Meijere, Java; C. javana de Meijere, Java; C. pusilla de 

 Meijere, Java; C. rugulosa de Meijere, Java. 



I have examined both European species. 



Stegana Meigen. 1830. Syst. Beschr., 6, 79. 



Phortica Schiner. 1862. Wien. ent. Monatsschr., 6, 433. 

 Amiola Loew. 1862. Berlin, ent. Zeit., 6. 

 Eostegana Hendel. 1913. Deutsch. ent. Zeit., 390. 

 Orthostegana Hendel. 1913. Deutsch. ent. Zeit. 



Arista plumose; three large orbitals, placed high up, uppermost one nearer to verticals 

 than to lowest (proclinate) orbital; postverticals small; eyes nearly bare; vibrissse and 

 ocellars present; one humeral; one presutural; two dorsocentrals; prescutellars present; 

 two scutellar pairs; mesopleurse bare; preapicals on all tibise; discal and second basal 

 cells separated; costa reaches fourth vein; anal cell and anal vein present. 



I have ventured to unite the two long-recognized genera Stegana and 

 Phortica. They are usually (e. g., by Schiner, by Williston, and by Olden- 

 berg) stated to differ in that Stegana has wings bent down at the base and 

 has the third and fourth veins strongly convergent, while in Phortica the 

 wings are straight and the third and fourth veins are parallel or nearly so. 

 But these two characters are not always associated. Hendel has described 

 two new genera: Eostegana, with bent-down wings but first posterior cell 

 not narrowed; Orthostegana, with wings straight but first posterior cell 

 narrowed. The narrowing of the first posterior cell is a character that shows 

 variations from species to species, so that it is hard to know where to draw 

 the line between "narrowed" and "not narrowed." The bending down 

 of the wings is still more elusive, at least in pinned material. I am con- 

 vinced, from an examination of many specimens, including the types, that 

 Phortica vittata Coquillett is based on specimens of typical Stegana in which 

 the wings are straight. I have seen specimens in which one wing was 

 straight, the other bent down. Specimens of Phortica humeralis Loew 

 sometimes have the wings bent down. These two genera are perhaps dis- 



