The Origin of the Markings of Caterpillars. [97 



give a definite reply on this point, but un- 

 fortunately we are not acquainted with any of 

 the young forms, so that we can but presume 

 that some of them at least would show only in the 

 first stage the simple subdorsal line without eye- 

 spots ; that in the second stage the primary pairs 

 of eye-spots would be formed on the fourth and 

 fifth segments, whilst the transference of these 

 spots to the remaining segments would take place 

 in the last stage. 



The foregoing assumption is based immediately 

 on the ontogeny of Elpenor and Porcellus ; it is 

 supported by the considerable size attained by the 

 eye-spots in many species of the third group, 

 and would receive additional confirmation by 

 observations on the Indian C. Celerio, supposing 

 that Horsfield's statements do not arise from a 

 confusion of species. This skilful observer, who 

 was the first to breed systematically a large 

 number of tropical larvae, has given a figure of the 

 Indian caterpillar of C. Celerio y according to which 

 this species possesses eye-spots on all the segments 

 from the fourth to the tenth. The European 

 form of this same species has eye-spots only on 

 segments four and five, a fact which does not 

 appear to have been known to Horsfield, as no 

 mention of it is made in his notice of the Indian 

 species. If the caterpillar figured is really that 

 of Celerio, which I consider to be by no means 

 improbable, not only is it thus shown that in the 



