Quagga and BurchelVs Zebra. 51 7 



example. In tlie latter, as in the Paris example, the pale 

 intervening areas are distinct upon the withers ; but behind 

 the withers in the Paris example they die out, the flanks 

 being only indistinctly striped and the hind-quarters practi- 

 cally unstriped. In Lorenzi^ on the other hand, the inter- 

 vening spaces persist in such a manner as to leave no doubt 

 that both in pattern and posterior extension the stripes were, 

 to all intents and purposes, like those of typical Burchell's 

 zebra {E, quagga BurchelU), tiie so-called " saddle ^' (" selle " 

 of Trouessart), characteristic of that animal and the more 

 northern forms related to it, being quite evident *. Herein 

 lies the chief difference between the Paris and Vienna 

 specimens. To the type of Greyi^ on the contrary, the Paris 

 specimen shows a close resemblance in the obliteration of the 

 stripes on the body and hind-quarters. 



Further evidence of likeness between the three specimens 

 above discussed, and especially between the Vienna and Paris 

 animals, is supplied by Dr. Trouessart's statement that the 

 latter has the appearance of a chestnut horse banded with 

 white, the stripes being brown and the intervening areas 

 whitish. In the typical quagga and E. q. iJanielli, on the 

 other hand, the stripes were black and the intervening areas 

 chestnut. 



It will be evident from what has been said that the Paris 

 specimen is to a great extent intermediate in its characters 

 between the types of Lore.nzi and Greyi. This fact may be 

 used as an argument in favour of the view held in 1904 by 

 Mr. Lydekkerf, that all the genuine quaggas belonged to a 

 single species very variable in the degree of development of 

 its stripes, but not resolvable into geographical races or sub- 

 species ; and also in support of tlie opinion, maintained by 

 myself, that there were several local forms of this animal, the 

 assumption of the probable existence of intermediates justi- 

 fying the view that only a subspecific value should be 

 assigned to the differences between them. Whatever con- 

 clusion be formed with regard to this matter, the cliief interest 

 of Dr. Trouessart's paper upon the Paris specimen lies in the 

 fact that it has proved the former existence of a quagga 



* As I have already pointed out, the pattern of the stripes on the body 

 and hind-quarters of the type of Lorenzi ati'ords convincinj^ evidence of 

 the nearness of the affinity between this quagga and typical BurchelU. 

 Dr. Lorenz also was forcibly struck by the siniihirity between the two 

 animals in this respect. The resemblance between them makes it im- 

 possible to draw up a logical definition of " quaggas " as distinct from 

 '• Burchell's zebras.' 



t P. Z. S. 1904, i. pp. 426-431. 



