Greek and Latin in Biological Nomenclature 39 



NOUN SUFFIXES 



Agent 



-TOR (-SOR), m., -TRIX, f., primary or secondary verbal; rarely 

 denominative: da-tor, giver; ora-tor, speaker; ton-sor, 

 barber; via-tor, traveler; impera-tor, commander; 

 pise a- tor, fisherman 



-ES (stem -T, -IT, -ET), m. or f., primary: ped-es, walker; com- 

 es, companion ; tram-es, pathway 



-o (stem -ON) m., primary: combib-o, pot-companion; ger-o, 

 porter 



-ARIUS, m., primary or secondary, verbal or denominative: 

 ferr-arius, smith ; pisc-arius, fishmonger 



Means or Instrument 



-BULUM, n., primary or secondary, usually verbal : pa-bulum, 

 fodder; vesti-bulum, court 



-CULUM (-CLUM), -CRUM, n., primary or secondary, usually 

 verbal : sar-culum, hoe ; vin-clum, fetter ; ful-crum, sup- 

 port 



-BRUM, n., -BRA, f., primary or secondary, verbal: cri-brum, 

 sieve; fla-brwn, blast; tere-bra, borer 



-TRUM_. n., -TRA, f., -TER, m., primary or secondary: plaus-trum, 

 wagon ; mulc-tra, milk-pail ; cul-ter, knife 



-MEN, -MENTUM, n., primary or secondary, usually verbal: 

 teg-men, cover; funda-mentum, foundation 



Action or Result 



-OR, -is, m., -ES, f., -us, n. (stem -OR, -ER), primary: cal-or, 



heat ; lab-or, toil ; cin-is, ashes ; pulv-is, dust ; caed-es, 



a setting-down ; nub-es, cloud ; fun-us, burial ; on-us, 



burden 

 -TUS (-sus), mostly masculine, primary: fruc-tus, fruit; pas- 



tus, food; vic-tns, food; sen-sus, perception 

 -10 (stem -ION), f., secondary verbal, rarely primary: obsid-io, 



siege; reg-io, district 

 -TIO (-sio), (stem -TION), primary or secondary verbal: auc- 



tio, increase; huma-tio, burial; conver-sio, alteration 

 TURA (-SURA), primary or secondary verbal: tex-tura, web; 



commis-siira, joint 



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