First Eremochcetous Dipteron with Vestigial Wings. 323 



XfiT. — I'lie First Kremocli<clous Dipteron loilh Vestigial 

 Wings. V>y Piof. .M. Hkzzi, Tiuiii, It.ily. 



In my pnper on tho reduction and di.sai)[)eaiaiice ot" the wiiiff.s 

 in ilixi Dipteiu* I have pointed out the tact tliat this pheno- 

 menon is very rare in the suborder of the Diptera Bracliycera. 

 In a total of 384 cases observed (adding to my list on 

 pp. 1()5-182 those appended at end of the present paper), we 

 liave the toliowing figures : — 



Suborder Orlhorrhapha Neniatocera . . . 140 

 Orthorrhapha lirachycera ... 12 

 Cyclorrhapha Athericera .... 232 



And even the few species of tho Brachycera are re)ircsented 

 exclusively by some Einpididai and Dolichopodidje, both 

 ])eloiiging to the same group of the Orthogenya (Microj)hona). 

 All the other groups of the suborder are exempt (the Hypo- 

 cera being placed with tlie CJyclorrhapha) ; in the Energopoda 

 there are a itw Asilidse in which the wings show a tendency 

 towards reduction both in the size and in venation, but these 

 cases seem to be unworthy of consideration. It was therefore 

 with the greatest interest that I received, some months ago, 

 through the courtesy of Dr. Charles P. Alexander, Urbana, 

 111., U.S.A., two specimens of a subupterous Dipteron from 

 Tasmania, sent to him with a collection of Tipulidai. On a 

 superficial examination the insect is indeed \e\y like a wingless 

 female of the genus Tipida, of which there are numerous 

 cases in all parts of the world. It was first believed to be a 

 IJibionid or a lihagionid (Leptid), but at once I recognized 

 in it a Chironi} zid, judging it to be a wingless female of some 

 Australian species of Chiromyza or of Metvpnuia. 



In the meantime, a paper f by Mr. G. H. Hardy, of the 

 Australian Museum, Sydney, N.S.W., has appeared, in 

 which an insect almost identical is described as a new genus 

 and species under the name of Borcoides kulndatus, with the 

 intere:?ting notice that it has been preserved for many years 

 in the Melhounie Museum under the MS. w^vwq o^ Boreovnjia 

 subulata, Walker. The genus is considered to be nearly 

 allied to Chiromyza. 1 am also of the same opinion. Tlie 

 atrophied mouth-parts, the form of the head and antenna), the 



* " Riduzione e scomparsa delle ali nej^U Insetti Uitteri," Eivista di 

 Sci. Nat. "Natura," Milano, vol. vii. 191(J, pp. 85-182, 11 figs. 



t " A Revision of the Cliiromyziiii (Diptera," IVoc. Linn, fcjoc. N.S.W. 

 xlv. 1920, pp. 532-542, j.l-s. xxi.x., xxx. 



