44 SEAT OF THE SENSE OF SMELL 



found that in some cases (though by no means in all), 

 insects which had been deprived of their antennae still 

 appeared to possess the sense of smell. But if, as we 

 have, I think, good reason to suppose, the power of 

 smell resides partly in the palpi, this would naturally 

 be the case. 



He also tested a beetle, Silpha thoraciea, with oil of 

 rosemary and assafoetida. It showed its perception 

 by a movement in half a second to a second in the 

 case of the oil of rosemary, and rather longer — one 

 second to two seconds — in the case of the assafoetida. 

 He then deprived it of its antennae, after which 

 it showed its perception of the oil of rosemary in 

 three seconds on an average of eleven trials ; while in 

 no case did it show any indication of perceiving the 

 assafoetida even in sixty seconds. 



This would seem to indicate a further complication — 

 not only that both the antennae and the palpi may 

 possess the sense of smell, but also that certain odours 

 may be perceived by the former, and others by the latter. 



Graber questions some of the experiments which 

 seemed to me * to demonstrate the existence of a sense 

 of smell in ants.f 



* " Ants, Bees, and "Wasps." 



t He says, "Da Lubbock nocli liinzufiigt, dass keiner, der das 

 Benehmen der Ameisen unter dieseu Umstandeu beobachten wiirde, 

 den geriDgsten Zweifel an ihrem Geruchsvermogen haben konnte, 

 wahlte ich auch diese Methode, um zu erforsehen, wie sich etwa der 

 Fiihler beraubte Ameisen vcrhalten wurden. Ich war nicht wenig 

 uberrascht zu finden, dass auch diese (es haudelt sich um Formica 

 rufa) vor dem Kiechobjekt umkehrten. Um ganz sicher zu gehen, 

 versuchte ich's aber noch mit dem gleichen Arrangement aber mit 

 Weglassung des Biechstoffes, und siche da ! sie kehrten auch jetzt noch 

 um ! Bei genauerer Beobachtung der von einer Ameise vom Anfaug 

 an auf dem Papiersteg zuriickgelegten Strecke stellte sich auch bald 



