88  SMITHSONIAN    MISCELLANEOUS    COLLECTIONS  VOL.    96 
The  genitalia  of  Occanthus  have  been  described  by  Walker  (1922), 
who  shows  that  they  differ  from  those  of  Gryllus  only  in  a  few  unim- 
portant details.  The  spermatophore  of  Oecantlius  pelluceus  is  de- 
scribed and  figured  by  Boldyrev  (1913a),  Gerhardt  (1914),  and 
Hohorst  (1937).  It  resembles  the  spermatophore  of  Gryllus  except  in 
the  small  size  of  the  "  attachment  plate  "  and  in  the  greater  thickness 
of  the  duct,  but  Gerhardt  and  Hohorst  show  that  the  plate  in  the  case 
of  Oecanthus  does  not  enter  the  genital  chamber  of  the  female,  and 
that  the  spermatophore  is  held  in  place  entirely  by  the  duct,  which, 
according  to  Hohorst,  is  armed  near  its  tip  with  a  small  brush  of 
short  bristles.  While  the  spermatophore  lies  within  the  phallus  the 
duct  is  recurved  above  the  ampulla  as  in  Gryllus,  but  after  its  ejection 
and  insertion  in  the  female  the  duct  is  straightened  out,  as  shown  in 
figures  by  Gerhardt  and  Hohorst. 
A  quite  different  type  of  phallic  structure  occurs  in  Neniobius.  but 
it  is  one  clearly  derived  from  that  of  Gryllus  and  Gryllodes.  In 
Nemobius  fasciatus  (fig.  36)  the  epiphallic  margin  of  the  dorsal 
phallic  lobe  (F,  e)  has  been  carried  so  far  back,  and  the  ventral  lobe 
(zd)  set  so  far  forward,  that  the  dorsal  cavity  (dc)  is  flattened  out 
until  it  appears  as  a  shallow  depression  on  the  ventral  side  of  the 
dorsal  lobe  (C,  F,  dc),  and  the  endophallic  cavity  (Enph)  comes  to 
lie  in  front  of  it  above  the  ventral  lobe  (vl).  The  two  cavities  are 
separated  by  a  U-shaped  fold  (g),  which  represents  the  originally 
posterior  lip  of  the  dorsal  cavity.  The  dorsal  cavity  of  the  phallus 
has  thus  undergone  a  complete  inversion  in  Ncinohius,  since  it  now 
occupies  the  ventral  surface  of  the  dorsal  lobe,  where  it  is  entirely 
exposed  on  removal  of  the  ninth  sternum  from  beneath  it  (C,  dc). 
When  the  phallus  of  Nemobius  is  viewed  from  above  (fig.  36  B), 
the  epiphallic  surface  of  the  dorsal  lobe  is  seen  to  have  the  form  of 
a  long  truncate  cone  with  a  large  V-shaped  sclerite  {in)  in  its  basal 
half,  and  a  weakly  sclerotized,  deeply  emarginate  distal  area  {e).  At 
the  sides  are  two  elongate  sclerites  (p)  ending  distally  in  two  pairs 
of  strong  lateral  processes  {r,  s) ,  and  a  pair  of  ventrally  convergent 
mesal  processes.  The  ventral  surface  of  the  dorsal  lobe  (C),  as  seen 
by  removal  of  the  ninth  sternum,  presents  a  long,  shallow  median 
depression  {dc),  which  is  the  inverted  dorsal  cavity,  limited  anteriorly 
by  the  U-shaped  fold  {g)  above  noted,  which  corresponds  with  the 
ventral  lip  of  the  dorsal  cavity  in  Ceuthophilus,  Gryllus,  and  Gryllodes 
(figs.  30  D,  32  E,  35  J,  g).  An  elongate  plate  on  the  floor  of  the 
cavity  tapers  distally  into  a  groove  between  the  halves  of  a  soft, 
globular,  median  lobe  {v).  Since  the  duct  of  the  spermatophore  passes 
through  this  groove,  the  cleft  lobe  evidently  represents  the  virgalike 
