10 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 



KEY TO SUBFAMILIES OF ENOPLIDAE 



1. (10) Buccal capsule surrounded by the esophageal muscula- 



ture ; absent in certain genera. 



2. (3) The cephalic armor (i.e., the chitinized part of the ex- 



ternal cephalic cuticle) present. Three well-devel- 

 oped lips present. The buccal organs complicated, 

 consisting of three unequally developed jaws (re- 

 duced in Saveljevia) with a cuticular ring around 

 them, homologous to the cephalic capsule of other 

 forms ; onchi in some genera Enoplinae 



3. (2) The cephalic armor absent (in Phanoderma there is 



present, besides the true cephalic capsule, which is 

 always absent in Enoplinae, a longitudinally striated 

 cervical armor). Lips mostly absent. 



4. (s) Esophageal contour straight; the cephalic circle always 



single, composed of 10 setae or papillae Leptosomatinae 



5. (4) Esophageal contour crenated. 



6. (9) Spear absent; buccal capsule absent or very small. 



7. (8) Esophagus broadly incised posteriorly. Body mostly 



elongate, in some species very thin. Spicules short, 



preanal tube absent Oxystominae 



8. (7) Esophagus rounded posteriorly. Body mostly moder- 



ately elongate. Spicules long, preanal tube present 



in most forms Phanodermatinae 



9. (6) Spear present, long and thin, followed by a modified 



part of the esophageal tube loosely articulated with 



it Thoracostomopsinae 



ID. (i) The buccal capsule free (as least for a part), generally 

 with strong chitinized walls (except when reduced 

 as in Enchelidium) . 



11. (14) The buccal capsule closed by six mobile lips, its walls 



strongly chitinized, mostly bearing three immobile 

 onchi. 



12. (13) The walls of the capsule without clefts, therefore 



completely immobile. Spicules very dissimilar; one 



preanal tube or none Oncholaiminae 



13. (12) The walls of the buccal capsule with clefts, therefore 



somewhat mobile. Spicules thin, curved; two pre- 

 anal tubes Eurystominae (Ditlevsenella) 



14. (11) Three lips or none, buccal capsule broadly open. 



15. (16) Buccal capsule conoid with chitinous folds in the walls, 



teeth short Rhabdodemaniinae 



16. (15) Buccal capsule cup- shaped with spear-shaped tooth or 



teeth, or reduced to form a narrow chitinous tube. 



17. (18) Spicules curved, preanal tubes two; one tooth Eurystominae 



18. (17) Spicules elongated, no preanal tubes; one tooth, three 



teeth, or none Enchelidiinae 



