120 THE ENTOMOLOGIST. 



in Mthalion, but are a trifle shorter, more oviform, and more 

 distinctly separated from the proportionately longer and thinner 

 apical part, which con^&ts, as in^Ulopa, pfi numerous regularly 

 disposed segments. In Paropia scanica aha Acocephalus striatus 

 the sensory parts are slender and clearly., divided into six seg- 

 ments ; the rest of the flagellum is, in Acocephalus, almost 

 identical vfitrrLedra ; i^ Paropia it is longer, and segmented 

 regularly up to the apex. j 



In sexually immature forms (of which I have examined Ledra 

 and two of our indigenous Jassinae (sens, lat.) ), the antennae 

 do not deviate in structure to any essential extent from the 

 sexually matured. 



(To be continued.) 



DIPHXLETISM IN THE LEPIDOPTERA. 

 By a. Radcliffe Gkote, A.M. 



In letters to me Dr. T. A. Chapman proposes to call in the 

 future my numbered vein " IX." of the fore wings in the 

 Papilionides, " Grote's vein," since upon the presence of this 

 vein I have based a separation of the swallow-tail group from 

 the rest of the butterflies, and because also a name which is 

 noncommittal is useful when applied to an organ the homologies 

 of which are doubtful, as in the present instance. At the same 

 time. Dr. Chapman suggests that this vein may be not the third 

 anal, which from its position it might seem to be, but the fourth 

 in the series of internal veins on the primary wing. 



While in my original papers I disputed the homology of 

 "Grote's vein" with the "fork" of the Hesperiades, which 

 Prof. Comstock regards as the remains of the third pupal vein, 

 it did not occur to me that it might be the fourth, since there 

 seemed no space between "Grote's vein" at base and the 

 second anal to admit of an intercalary longitudinal vein. In my 

 paper on the " Descent of the Pierids," I have given a tentative 

 sketch of a genealogical tree of the Hesperiad phylum, deriving 

 it from an ancestry in which four anal veins on either wing were 

 present, and which would conform with ontogeny. But this four- 

 veined state, now observed in the pupal wing, represents in reality 

 a common ground upon which the existing types of lepidopterous 

 wings have arisen by specialization through reduction. That, in 

 the Hesperiades, the fourth vein appears in the pupa, is no proof 

 that the Papilionides have the same origin, since we have to do 

 with a general character shown also by moths in ontogeny. It 

 seems to indicate rather that the diurnal branch referred to sprang 

 directly from a hypothetical Tineid-like ancestry, as I have sug- 

 gested in the paper already mentioned. Thus the Hesperiades, 



