259 



ON SOME RECENT ATTEMPTS TO CLASSIFY THE 

 COLEOPTERA IN ACCORDANCE WITH THEIR 

 PHYLOGENY. 



By C. J. Gahan, M.A. 



Published by Permission of the Trustees of the British Museum.) 

 (Continued from p. 248.) 



Ganglbauer's Classification. 



Ganglbauer divides the Coleoptera into suborders characterised 

 as follows : — 



I. Suborder Adephaga. — Wing-venation of the first type. 

 Testes simple, tubular, Male genital apparatus with only one 

 pair of accessory glands (ectadenia). Ovaries with alternating 

 nutritive and egg-chambers. Four Malpighian vessels. Larvae 

 orthognathous ; campodeiform or only slightly departing from 

 that form ; with two-jointed tarsi. 



One family-series. Families : Carabidse, Dytiscidae, Haliplidae, 

 Gyrinidae, Rhysodidse, Cupedidae, Paussidae. 



II. Suborder Polyphaga. — Wing-venation of the second or 

 third types. Testes compound, consisting of follicles. Male genital 

 apparatus with one or more pairs of accessory glands. Ovaries 

 with single terminal nutritive chamber. Four or six Malpighian 

 vessels. Larvae with long or short legs, having one-jointed tarsi, 

 or without legs ; campodeiform or eruciform ; orthognathous, 

 hypognathous, or pseudorthognathous. 



1. Family-Series Staphylinoidea. 

 Wing-venation of the second type, in some forms so reduced 



that the type cannot be recognized. Gular sutures not confluent. 

 Pleural sutures of prothorax distinct.* Antennae simple, or 

 with the terminal joints thickened to form an unflattened club, 

 occasionally irregular. Tarsi with a variable number of joints. 

 Testicular follicles sessile. Male genital apparatus with two 

 pairs of accessory glands (ectadenia and mesadenia). Four 

 Malpighian vessels. Larvae campodeiform, or not widely deriva- 

 tive therefrom, never grub-like or eruciform. 



Families : Staphylinidae, Pselaphidae, Scydmaenidae, Silphidae, 

 Clambidae, Leptinidae, Platypsyllidae, Aphaenocephalidae, Corylo- 

 phidae, Sphasriidae, Trichopterygidae, Hydroscaphidae, Scaphi- 

 diidae, Histeridae. 



2. Family- Series Diversicornia. 

 Wing-venation of the third type, or approximating towards 



the second type, in some forms very reduced, and then without 



■'- For pleural we should read pleuro-sternal, both here and in the 

 following groups, in order that this character may be correct. — C. J. G. 



u 2 



