8 



MORPHOLOGY 



however, nowhere prevails : the portion of tube ^J"-*£^ ^ 



base, whether short or 



of uniform calibre. The staminal insertion in P. pectmata 



ment 



an? J Tenulrostns is at the° base of a transverse indentation of the tube, but this arrange 



occurs nowhere else, and is absent from the very nearly allied P. pyrarnidata. 

 The tube in Siphonantile and in Orthorrhynch^e is straight, but in many Rhyncho- 

 lophe and in most Bidehtat* it is curved, the concavity being anterior. In Anodont^ 

 it bWf frequently angularly bent either within the tube or opposite the hmbus of 



the calyx In some cases this infraction is absent, and the tube may be straight (P 

 Kingii) or incurved (P. pycnantha). The amount of infraction varies somewhat, but the 

 situation of the angle is constant for individual species. 



The lip is always 3-lobed, and usually is rounded in front, very rarely narrowing 

 gradually from a wide base. In Anodont^e and Bidbntat^b the lip is stipitate, the base 

 bem" erect and 2-cristate above, and the lobes usually too small to envelope the hood in 

 the bud. Even when sufficiently large to do so, they are early deflected and permit the 

 hood to appear. In Siphonanthje and Orthorrhyncm the lip is sessile, the lateral lobes 



lapping in the bud and quite covering the hood, while the base is narrow and 



uncreated. In Rhyncholoph.e the lip may be stipitate with a wide base and with distinct 



crests (P. Colleltii, zeylanica), with a narrow base and then with only rudimentary folds 

 near the base of the lobes (P. Clarkei, Prainiana), or without folds (P. lachnoglossa) ; 

 or it may be sessile (P. carnosa, gruina) and without folds. The lobes are usually 

 rounded, with the margin entire, or with the midlobe (P. siphonantha), or all three (P. 

 longijlora) truncate, but are sometimes ovate (P. corymbosa), or obovate (P. Perrottetii). 

 When rounded the midlobe is as a rule somewhat narrower than the lateral lobes; 



when ovate or obovate the lobes are nearly always equal in size. In a few cases (P. 

 Kingii, fragilis) the lateral lobes are narrower than the central; in these instances the 

 midlobe is rounded while the lateral lobes are ovate. The margin is often ciliate, a 

 character which is extremely constant in individual species. 



The hood consists of two distinct portions — a basal continuous with the tube and an 



upper which encloses the anthers. The basal part is always of the same consistence as 



the tube and generally of the same width, is always erect and generally of the same 



length as the upper portion, though it may be distinctly longer (P. schizorrhyncha), or 



may, on the other hand, be almost absent (P. excelsa). The margin is usually distinctly 



thickened, sometimes sinuate (P. longijlora, furfuracea), sometimes toothed (P. siphonantha, 



odonhphora), and sometimes replicate (P. Oederi, pycnantha). The upper part or hood 



proper is, except in Anodont.e, of firmer consistence, and is generally a little wider than 



the basal portion. In ANODONTiE it is erect and continuous with the base ; in Bidentat^, 



most Rhyncholopb^e, and in Siphonantkle it is curved; while in Orthorrhynch^e it is 



angularly bent. The margin in Anodont^e and in some Rblyncholophje is entire or is 



toothed only at the lower angle (P. Regeliana) ; in Bidentatje it is toothed immediately 



below the apex; while in most Rhyncholophje and in all save one species of the 



LONGIROSTRES it is prolonged at the apex into a distinct beak. The beak, like the tube, 



is late of development, and though discernible in the bud, it is impossible to say from its 



condition there whether it will remain rudimentary (P. globifera), become short and thick 



(P. alaschanica), slender and falcate (P. gruina), straight (P.flexuosa), flexuose (P. pedinata), 



or circinnate (P. siphonantha). Neither the absolute length of beak nor the relative 



lengths of beak and lip or of beak and hood afford reliable characters in diagnosis. 



The apex of the beak may be entire, emarginate, or deeply bifid, and this condition, which 



I 



