

.-ESTIVATION, 



9 



is 



dent in the very youngest bud, is of high diagnostic value. The estivation of the 



as to the structural affinities of species. Owino- 

 to the lateness of development of the beak, young buds afford less information than mi<*t 



beak appears to afford important evidence 



be 



pected. But the interpretation of the cond 



dental circumstances 



In the Siphon an mm the flo 



is materially assisted by 



bracts, if they open at all, open later than those 



in the axils cf th 



flowers are not, however, 



higher 



up 



These lower 



lower 

 pened 



sarily undeveloped; on the contrary, the beak elongate 



till it attains its full size and the tube also elongates and lies in zig-zag folds imprisoned 

 in the lower portion of the calyx-cup. Only careful study in the field can determine 



hether we have here a mere case of 



togamy 



rn 



fh 



phological aspect of the 



trifugal 



biological, and 



SO fi 



opening or a condition akin to clcis- 



is, however, as interesting as the 



gement 



Axis of beak parallel to median line 



as the beak is concerned, admits of 



following defi 



th direction reversed, since the dorsal entire 



aspect of the beak is apposed to the lip, and the apex is directed to the throat of the 

 corolla. It may be expressed by the notation 



beak, the arrow L 



present 



ti 



B 



L 

 In all 



(the arrow R standing for the 



an 



opposite 



d 



prevails. It is seen in its simplest form in species where beak and lip are of 



length (P. gracilis, Jlexuosa), but 



(P. pectinata, pyramidattz), and may be defined as f 



is equally evident in species 



with long fl 



same 



beaks 



line of Up with direction the same 



to the lip, and the 



Axis of beak parallel to media 



j 



the ventral divided aspect of the beak is 



ap 



is 



directed to the margin of its 



ddle lobe 



1 



apposed 



notation 



here would b 



R 

 L 



as 



before rep 



the beak and L the 



P 



It is not easy to 



imag 



a condition intermediate bet 



two such 



gements, but it is interesting to find that in most of the beaked Rhy 



opp 



cond 



exists of which 



the two arrangements 



a 



already described might be 



derivates. It may be stated thus x—Azis of beak perpendicular to median line 



qually well 



to ith 



direction 



t 



ight 



igles, since the ventral divided aspect of the beak is directed towards 



the throat of the coroll 



d 



apex 



is 



pposed to the middle of the 



P 



This 



di 



tion may be symbolized as j 



(where R is again the beak, L the \\ 



l>) 



) 



The correlation of beak and tube deserves notice. In those species where the beak 



pronounced the 



tube tends to become of uniform calibre 



those species wh 



the corolla tube is infundibuliform there is no beak 



This is made more striking 



g owing 



to the fact that in the Siphonanth^, of which a tube with uniform calibre is highly 



characteristic, species which have 



short beak (P. macrantha, robusta) h 



widened tube, and i 

 which some memb 



the highly natural groups, so common 



amongst Rhy 



ightly 

 E. of 



possess 



wh 



others are without beaks, the beaked forms 



carnosa, gruina, polyphylla) have almost 



niform tubes, while the correspondin 



b 



beakl 



ess 



ones (P. zeylanica, 

 in this alteration 



Henry i, tenuisect 



nfundibulifcrm 



Tl 



lip also is involved 



; as the beak elongates an 

 ests disappear, and its lobes enlarge. Insid 



d th 



tube constricts its base 



the tube in front it is 



shortens, its 

 al to find in 



ort-beaked species a number 



of long hah 



th 



m 



a y 



be 



onfined to the lo 



par 



of the tube (P. zeyl 



or 



extend to the base of the lip (P. rex). This condition also 



tends to disappear when the corolla tube 



is contracted, as it is in long beaked 



pec 



Amn. Roy. Bot. Gard. Calcutta, Vol III. 



