Anther single— 
Tribe 1.— MALAXIDEAE. Pollinia 2 or 4, waxy, without appendages, free or rarely 
attached by their bases to a ball of translucent matter. 
Tribe Il.—EPIDENDREAE. Pollinia 4 or 8, waxy, attached by their bases or by 
their backs to а single or double granular or translucent appendage, 
but without any gland derived from the stigma. 
Tribe IIl.—VANDEAE. Pollinia 2, 4, or rarely 8, attached singly or in pairs or in 
fours by a caudicle (rarely by two caudicles) to a viscid gland derived 
from the stigma (in a few doubtfully from the stigma). 
Tribe IV.—LISTEREAE. Anther terminal (dorsal in Epipactis), pollinia two or four, 
cohering by their sides, without gland or caudicle (a rudimentary 
caudicle in Epipogum); pollen powdery, granular or in small masses, 
never waxy. 
Tribe V.—GOODYEREAE. Ап ег posticous, vertical but inverted ; pollinia two, or four 
in two pairs, attached to a single gland either by one or by two caudi- 
cles, or without a caudicle ; pollen granular or sectile. 
Anthers two, each with only one perfect cell— 
Tribe VI.—OPHRYDEAE. Anther-cells sessile on the column, anticous or posticous, 
discrete or contiguous but always distinct, parallel or diverging, often 
with tubes from their bases, each with a staminode on its outer surface ; 
pollinia two, rarely four, usually with caudicles and always with 
glands; the glands naked or enclosed in the same pouch or in two 
distinct pouches, Stigmas two, distinct or conjoined below the anther- 
cells ; pollen granular or sectile, not waxy. 
Anthers two, each perfect— | 
Tribe VII.—CYPRIPEDIEAE. Perfect anthers two, one on each side of the conjoined 
stigmas. *Staminode single, large, fleshy; lip large, saccate. 
