64 



MEMOIRS OF THE NATIONAL ACADEMY OF SCIENCES. 



rti^p 



riG. 10 Head of 



pu]taof Grac'ilaria; ci, 

 clylnMls: I, laUruin. 



ami also those with riiiK'tional or vestijii;il maxillary palpi, sucli as Tiiieidie, GracilaiiidiE, 

 Elachistidie, etc. 



It is evident that the elassifleatiou ol' the Tiiieiiia will have to he entirely recast. Instead of 

 placing the Tineida', with their broad wings and generalized venation, at the head oltlie Tineina, 

 as done in our catalojiues and general works, they should go to the base of the series, not far 

 from the Microi)terygi(la'. On looking over the venation of the Tineida' rei)re- 

 sented on Spuler's IM. XW'l, it is evident that the very narrow-winged genera 

 such as Coleophora, Ornix, Lithocolletis, Nepticula, (Jelechia, Cemiostoina. and 

 CEcophora, are highly modified recent forms when compared with Tinea and 

 Blabophanes, as well as the Adelidiu (Adela, fig. 7), Nematois (fig. 8), and Choreu. 

 tida' (Sinnvthis, fig. 0, larva, and Choreutis), and justify Chapman in associating 

 them with the Pyraloids in his group of Pitpcc ubtecta: 



The pupa of Gracilaria (flg. 10) and of Bucculatrix (tig. 11) shows the eye- 

 collar, the paraclypeal tubercles, as well as the labial palpi. On the other 

 hand, the pupa of the pyraloid genus Cryptolechia (figs. 22, 23, C. quercicelln, C. 

 schh(fiiiiicUa) shows no traces of the maxillary palpi (eye-collar). 



Famil!/ Proiloxlda'. — Uaving already discussed the chief characteristics of 

 the Palsolepidoptera, represcuted by the family Micropterygidie, we may next 

 call attention to tlu' most primitive of the Neolepidoptera. These we believe 

 to be the very remarkable genera Tegeticula (Pronuba) and I'rodoxus, repre- 

 senting the family Prodoxida'. The structure of the imagines and their larval 

 and pupal forms have been described at length and figured by Dr. C. Y. IJiley,' 

 who has de.-<-ribed the egg as being very long, cylindrical, soft, and flexible: 

 the boring larva; as being either without abdominal legs, but with thoracic 

 ones (Tegeticula), or entirely apodous (Pronuba). Dr. IJiley gives a careful and 

 detailed account of the male and female pupa of Tegeticula- (Pronuba), but does 

 nat mention the "eye-ct)llar" or case of the end of the maxillary ]}a\\)i (figs. 12, 13, ww. p.)^ which 

 is very large, especially in Tegeticula, much more so than in the rest of the Tineina or in any 

 of the other Neolepidoptera. It is thus in a degree intermediate between that of the Neo- and 

 Paleolepido[)tera. The maxilla- {m.x:) are well developed, but there are no traces, so far as 1 can 



see, of the ''maxillary tentacles'' so greatly develojjed, 

 according to Eiley, in the imago; but the si)ecimens kindly 

 lent me by Dr. Riley for examination are 

 the cast shells, and further examiinition 

 and search for them should be made on 

 living or alcoholii- s|)ecimens. The labial 

 palpi [mx. p.) and the paraclypeal pieces, 

 as well as the eye-suture sei)arating the 

 •'glazed eye" from the rest of the eye, are 

 well developed. ^\bdominal segments 2-9 

 are free and armed with the enormous 

 dorsal spines well described and figured 

 by Riley. Figs. 11 and 15 represent the 

 cast i)upa skin of Trodoxus decipiens 

 Riley. 



The venation is almost exactly as in 

 Tineida?, but the structure of the maxilhe, 

 as des(;ribed and figured by Riley, jtrescnts an extraordinary feature, in which this family, and 

 es])ecially the present genus, differs from all the other insects. I refer to the remarkable 

 " maxillary tentacles." Riley thus describes them : 



TIr' male jiosscsaes no very iiiaiked iharacters, liiit tlie female is most anomalous; first, in possessing a jiair of 

 picbensile, spinous, nuixillar.v tentacles (fig. 16), found, so fur as wo now know, in no other genus of Lepidojitera. 



Fl(i. 11. — Puj>a "f Jluccutntrix 'ixtiiiqunintella ; ,1, oblitjiu-; J?, side view of 

 bead ; ch, coc-nou-buratcT; (\ sidti view of licad of C. canadensella. 



' Proc. Amer. Assoc. Adv. Sc, xxix, 1880. 



