8 



Piosmeae, in whicli the typical genus Diosma, though gq 

 dissimilar in habit, has many characters in common with 

 Acradpnia, its structural differences consist in its consp^cuQusly 

 6 lobed disk, its pentandrous flowers, and the total absence of 

 the albumen. 



It remains now to trace the nearest affinities of ^cradenia 

 among Xanthoxyleae. Pilocarpus, confined to tropical 

 America, possesses 4-5 stamens, it has moreover the calyx 

 merely minutely toothed, and the flowers are racemose or 

 spicate. But these notes become not singly but only col- 

 lectively of any value for generic separation as demonstrated 

 by Euodia, Boronia and Eriostemon. Among Australian 

 Xanthoxylese, Pagetia (F. Muell. fragm. phytogr. Austr. Y, 

 178-215), differs in the plurality of ovules, in the adnate 

 endocarp, and in the absence of albumen. Euodia (combined 

 ■with Melicope) has a tetrapetalous carolla, a long testa, a 

 copious albumen, foliaceous cotyledons, and a conspicuous 

 cylindrical radicle. Bosistoa (P. Muell. in Benth. fl. Austr. 

 I. 359,^ next to which Acradenia must be placed, though often 

 distinguished by pinnate leaves, shows these occasionally, if 

 not frequently, reduced to three leaflets, and it would not be 

 surprising if Acradenia like some Boroniae was found to vary 

 in this respect also. The petals of Bosistoa are valvate in 

 bud, a character of little moment in the mutual relation of 

 genera and even species, of Rutaceae ; the carpids are large, a 

 note of no generic importance though remarkable in this 

 instance, and the albumen is entirely wanting. 



To these distinctions so little importance can be attached 

 that the writer has been induced to locate Bosistoa sapindi- 

 formis as Acradenia Bosistoi (Australian vegetation, 

 indigenous and introduced, considered in its bearing to 

 territorial resources and settlement p. 29) under the Tasmanian 

 genus. Discoveries of new allied forms in the jungles of 

 East Australia may draw the connection between these plants 

 still closer. 



The two localities in which Acradenia Frankliniae is known, 

 are the vicinity of Port Davey and of Port Macquarie, but it 

 may be presumed that this noble and rare plant, so worthy to 

 bear the name of the high-minded Lady Franklin, occurs in 

 many of the interjacent deep forest ravines, where it is likely 

 to rise to a stately height. 



Melbourne Botanic Garden, 

 12th February, 1867. 



