S 
ZALACCA. 67 
ZALACCA  Reinw. 
Reinw. in lit. ad Mart. ex Mart. Hist. Nat. Palm. ii, 199, t. 118, 119, 128 
136 and 159, f. 2; Rumph. Herb. Amb. v, 113; Blume in. Schult. Syst. Veget. 
vii, 1333, Obs. 3; Blume, Rumphia, ii, 158; Endl, Gen. No. 1737; Meisn. Gen. 
354 (265); Wall. Pl. Asiat: Rar. ili, t. 229—224; Grifi. in (alc. Journal Nat. 
Hist. v, 8, and Palms Brit. Ind. 9, t. clxxv—elxxx; Mig. Fl. Ind. Bat. iu, 80; 
Beec. Malesia, iii, 63; Kurz, For. Fl. Brit. Burma ii, 511; Hook. f. Fl. Brit. Ind. 
vi, 472; Ridley, Mat. Fl. Mal. Penins. ii, 168; Salacca Reinw. in Syll. Plant. ii, 3. 
Cæspitose, almost stemless, spinous, dioecious palms, having large, pinnate, 
non-flagelliferous leaves. Leaflets lanceolate or linear-lanceolate, tricostulate, almost 
always sigmoid or at. least falcately acuminate. Spadices interfoliaceous, very 
dissimilar in the two sexes, enveloped in several incomplete membranous, mostly 
lacerated and perishable spathes. Male spadices branched, bearing several catkinlike, 
cylindrical spikes, composed of very approximate bracts (spathels) more or less connate 
by their margins. Male flowers two and equal at the axilla of every spathel, ovate, 
clavate or oblong, accompanied by special bracteoles usually hairy or woolly; the . 
calyx thinly membranous or hyaline,’ deeply 3-parted ; the corolla longer than the 
calyx, having a fleshy undivided base, and parted into 3 thinly cartilaginous lobes; 
stamens 6, inserted at the throat of the corolla; filaments short, subulate; anthers 
erect, basifixed; rudimentary ovary extremely minute, placed at the base of the 
tubular part of the corolla. Female spadix less branched than the male and with 
fewer spikes. Female spikes larger than the male ones, torulose or squarrose 
(Euzalacca and Eletodoxa), or small and composed of very few flowers (Letozalacea) ; 
the spathels at first connate by their margins, then separating, larger and less crowded 
than in the male spikes, each carrying a female flower accompanied by a neuter one 
(in the species of the Huzalacea and Eleiodora section), the female having two bracts, 
and the neuter only one ; the neuter flower is wanting in the species of the Lezozalacca 
section. Female flowers larger than the male ones, ovoid ; the calyx membranous. 
split into 3 parts; the corolla coriaceous, about as long as or slightly longer than the 
calyx, having a ventricose or urceolate entire base, and divided above into 3 valvate , 
segments; staminodes 6, inserted at the throat of the corolla; ovary distinctly 
3-celled, ovoid, strigose from being covered with scales prolonged into ascending or 
spreading spicule (Huzalacca), or less frequently flat and appressed (Letozalacca and 
Eleiodora). Fruit relatively large, _mono-trispermous, globose, turbinate or ovoid 
covered with reversed scales terminating in a rigid spiculiform upturned point 
(Euzalacca) or smooth and appressed (Lesozalacca and Eleiodora).. | Seed oblong, 
globular or obsoletely angular, enveloped by a fleshy, often abundant and very acid, 
integument of which an intrusion penetrates into the albumen from an apical pit; the 
surface of the nucleus smooth ; albumen homogeneous ; embryo basal. 
The genus is sharply divisible into 3 sections or subgenera. The more numerous 
and more characteristic representatives of ‘the genus (the Zuzalacca) are sharply 
separated from the others by the strigose ovary and the corresponding hispid subspinous 
fruit ; moreover in the species of this group the female flowers are accompanied by 
a neuter one and are crowded into cylindrical spikes. In the second group the female 
ANN. Roy. Bor. GARD., CALCUTTA, VOL, XII. 
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