Pammel — Anatomical Characters of Seeds of Leguminosae. 147 



occurs in other genera, as in Tonka bean, and in Hierochloe^ 

 a grass. 



Melilotus alba, Lam. 



Fl. VIII. f. 2. 



Testa and endosperm vary in thickness, average 75 ^. Mal- 

 pighian cells as long as the thickness of the endosperm and 

 remainder of testa. 



Malpighian. Cuticle wavy and well developed ; the cuticu- 

 larized layer below with small, conical projections, those of 

 two adjoining cells meeting at the middle lamella of the lateral 

 walls, giving the layer the appearance of consisting of cone- 

 like projections. These cones are also connected with the 

 small pore-canals. This cuticularized layer is highly refrac- 

 tive. The light line consists of a narrow but distinct refractive 

 zone below the conical layer. The refractive zone colors blue 

 with chlor-iodide of zinc. The whole upper part is, however, 

 more or less refractive. The remainder of tl:\e cell-wall 

 contains pigment and is colored blue with chlor-iodide of 

 zinc ; the cuticularized layer as well as the conical layer 

 colors blue. Small canals project into the walls, in some 

 cases extending beyond the light line. The chromatophores 

 are irregularly distributed in the cell-cavity, some near the 

 base, others in the center. 



Osteosclerid . Cells with a broad base and a small triangular 

 intercellular space above; longitudinal pore-canals in the 

 upper part of the cell, but these do not extend its entire 

 length. % 



Nutrient. This layer is much compressed; consists of 

 thin-walled cells, divided into two parts; cell-walls of lower 

 part thicker. Both layers contain pigment and tannin, the 

 upper more than the lower. Cell-walls consist of cellulose. 



Endosperm. The aleurone layer is quite distinct; the cells 

 are rectangular ; cell-walls made up of cellulose. The walls of 

 the remainder of the endosperm, except where it joins the 

 embryo, are thick, consisting of mucilaginous reserve cellu- 

 lose. Cells of the internal layer of the endosperm thick- 

 walled, elongated, containing some protein grains and fat. 



Embryo. Cells of the exterior walls of first row thickened ; 

 they are smaller than those below. All of the cells contain 



