ON Ti K P1BTILIZATION 





—I*" „f, ,11s h p~»1 .*-*"•" pi 



containing five or six 



■" l *:Xi ' KS« orifice a^ ..rejects somewhat beyond it (Plate V fig. 1} . 



,lrt,a : b ' S. 1 baJund micrepylar thickenings d.ffer in form from those in the rest 



Tfce C«'II» in bot " Ule , .„ „ r nn iv.*onal in place of flattened 



Within tllO epiucima fViinlrfimnors whfirft thft tisano 



WlthlB «»••!•""•» - nt8 basal and apical thickenings where the tissue 



,he large embryo** Thu > als p ^ y> figg _ The 



beeoni 



ba 



- much denaer and more coherent than it 

 hickoning consists o| many superimposed strata of cells, whn* are frequently 



"* ' : , , to 'form -,- « l<* distinctly defined lobe. (Plate V, fig. 7). The apical 



Sming i. very peculiar (Plate V, fig. 1). It forms a dense, broadly conical mass capping 

 t «x of the embryo-sac. The constituent cells are closely adapted to one another, are 



1Z <«lar or polygonal in outline, and contain relatively large, deeply-staining nuclei. 



fZLllv the cap is of great thickness, containing six or seven strata of cells (Plate V, fig. 1 V 



It tl HOff all round peripherally, and gradually subsides into the surrounding loose cellular 



the lat.ral surfaces of the embryo-sac. Under the influence of the reagents 



Zriov 1 in mounting pennanent ipeoimeM of ovules, and specially of entire ovules, the 



asuo covering 



iiu r nuOS 



coat frequently shrinks away from the epidermal one save at the base 



i 



|«ve a clear space betwc D them (Plate V, fig. 8). The walls of the cells do not show the 

 brownish tinge with picrocarmine which those of the epidermal stratum do, and the large 

 nuclei, save in the apical thickening, stain comparatively feebly. 



Within the general mass of nucellar parenchyma, as this stratum may be conveniently 



tanned and immediately around the embryo-sac, a certain number of small flattened cells 



■ 



ppearing fusiform in profile, are present (Plate V, fig. 1). These are most abundant, as a 



towards the apex of the nucellus. Immediately beneath the apical cap, and attached to 

 of its ron.stituent cells, is a large and peculiar cell of this type (Plate V, figs. 1, 2, 3, 4, 5, 6). 

 sectional view it appears as a curved spindle with the centre of the convex surface 



attached to the under surface of the apical cap, and the prominence of the concave one in 

 emit t with or in close relation to, the outer surface of the apex of the embryo-sac. This 



prominence is sometimes very marked, and where the embryo-sac has not shrunk too far 

 awa in the • >urse of preparation of the specimen, it often appears to depress its apex, while 

 the two pointed extremities of the spindle project free on either side as lateral horns (Plate 



'he entire body of the spindle is characterised by staining very deeply, 

 especially with logwood. 



The centre of the nucellus is occupied by a huge embryo-sac, with a delicate membran- 



wall a network of finely molecular protoplasm, and a large nucleolate nucleus (Plate 



y 



fig- 3). There do not, as a rule, appear to be any oosphere, synergidse, or antipodal 



cells. Only in one case have I been able to detect anything which might possibly be taken 

 to represent an oosphere and synergidaj, and in that the appearances were doubtful, and such 

 as could only correspond with elements of very abortive character. 



B.— < haracters of female flowers shortly after access of insects to the receptacles. 



^ The following show the measurements of a pedicellate flower a few days after the access 

 insects to the receptacle: — 



Total height from base of pedicel to summit of ovary 3*02 



eight from base to origin of the perianth 1-34 



H 



mm. 



Total height of ovary a. 



mm. 



79 mm. 



