OF FICUS KOXBrBGH!!. 



Breadth of ovary at level of upper edge of stylar b^rtion . 



??i i T T?:* ge ° f 8tykr ^^tosumu.itofomv' 1 m - 



Total length of style and stigma 7 ' , . mm 



Breadth of stigma 





0-6 



»5 mm 



1 V, 



oi; 



mm 

 mm 



These figures very dearly indicate the occurrence of rapid growth b the ovarie. «,d 

 the_ case of pedteellate flowers, in the ped.cels suhsequeut to the acce^i^! 



n 



project mi i 



penanth re ains its previous d.mens.ons, and the ovary consequently com,. , „ rojPCT „ 



and more from it. _ Even at the early stage represented hy the flowers of which ^-Tr 



ments have been given, the summit of the ovary rises high above the tip. of «h. „ 

 The origin of the style, just as in the case of the gall-flowers, appear. u> d 



ii 



f 





1 





g-wth m the ^upper part of the ovary, and iu some caaes the colour oi thc^ « 



parts of the atylea and the stigmata gradually changes i om -.madder to hrick Z 

 Beyond this neither styles nor stigmata show any change, and, with the x< ption of tin 

 basal portion of the style, which ultimately becomes softened, remain , *.* up • , tlu 

 period of maturation of the seeds and long after the outer coats of the o^ try have ui 1 

 mucoid degeneration. The walls of the ovary gradually thick n, the increase being 

 due to changes taking place in the two inner strata; the cells of tbe internal e] ^ 

 increasing in depth and in the thickness of their walls, and processes of cell multipli ation 

 occurring in the stratum of small cells lying external to them. The gen ral in reuse in he 

 bulk of the flowers even within a few days subsequent to access of insects to the r oeptael 

 is so considerable as to be very evident even to casual inspection by tl una dey< 

 The important phenomena are those manifesting themselves in the ovules. These, whi.i 

 removed from the ovaries a few days after access of insects to the receptacles and taminnl 



in water and uncovered so as to avoid flattening, give diameters of about 0*51 x0 14 mm 

 The secundine and nucellar epidermis show no important changes save those dependent u 

 extension, due to increased bulk of the deeper parts of the nucellus, but conspici us 

 changes soon set in in the nucellar parenchyma and embryo-sac. In the former there i 

 firstly, general growth around the sides of the embryo-sac, and secondly, special growth at its 

 base and apex. The cells of the loose tissue of the parenchyma begin to increase in size, the) 

 stain much more deeply than they did previously, and there is an obvious accumulation <>f 

 protoplasm within them (Plate V, fig. 9). Beneath the base of the embryo-sac the a< una 

 lation of cells becomes thicker and denser than it was before, and in some cases, at all ev nts 

 a peculiar local outgrowth takes place on its upper surface, giving rise to a prominent 

 circular mound of very small-celled tissue surrounding a central depression, and apparently 

 strongly cuticularised on the surface (Plate V, fig. 16). As it is developed, it pu lies up and 



invaginates the lower end of the embryo-sac. 



The walls of the cells of the apical cap generally become considerably thick tied, but do 

 not otherwise show any appreciable change. The special cell attached to its under surface, 



which appears as a deeply-stained spindle in section, on the other hand, undent 

 very remarkable development. It swells up centrally, and at the same time the peripheral 

 portions shoot out into large processes and become gradually separated by partitions from the 

 central dilatation (Plate V, figs. 11, 12). In sections it would appear as though we were 

 dealing with changes occurring in a simple spindle, but, judging from the appearance 

 present in some cases in entire or partially-dissected ovules, it appears to be probable 

 reality the cell oririnallv consists of a central more or less convex mass with radiant 



•> 



that 



Ann. Roy. Bot. Gard Calcutta, Vol. I. Appendix. 





