ANONACEEX OF BRITISH INDIA. 3 
account of the want of constancy in co-ordination already alluded to, For example, the 
Unonas of the section Dasymaschala have their outer petals united (their inner petals 
being also suppressed), although in no other respects do they differ from the Unonas 
with free petals) As regards direction there is some diversity, the petals of the species 
which have been grouped in the tribe Uvartee being spreading and almost rosaceous 
in their arrangement; those of the genus Artabotrys having saccate connivent bases, 
while the limbs are free and divergent. The imner petals of Oxymitra and of many 
other genera are, on the other hand, connivent; while the group of plants which 
have been collected under the genus Mitrephora have their spathulate petals coherent 
by their broad limbs to form an open mitre-like or Jantern-like corolla. In texture, 
the petals in this order have a tendency to be coriaceous, although a certain number 
-are membranous. As regards colour, an unusually large proportion are greenish-yellow 
or green. 
As a general rule, the stamens of Anonacee are numerous, and are arranged in 
spirals on a convex or flat torus, Some species have, however, only from 6 to 12 
stamens: these will be found in the genera Bocagea, Orophea and Mitrephora., 
In the stamens of this family the filament forms a very inconspicuous, and the 
connective a very conspicuous, part. In a large proportion of the species the connective 
is thick and is either oblong quadrate or cuneate, the truncate end projecting over 
the apex of the anther so as completely to conceal the anther-cells when looked at 
from above. ‘The elongated anther-cells are, in such stamens, attached vertically either 
at the sides or on the posterior surface of the connective. This type of stamen has 
been called the Uvarieoid. On the other hand, there are species in which the connective, 
instead of ‘being thick with a broad truncate apex, is compressed and. produced at the 
apex into a pointed or flattened appendage by which, when looked at from above, the 
apices of the anther-cells are not concealed. Stamens of this latter sort are usually 
associated with the anomalies in the petals which have already been referred to; and 
the plants possessing them have been grouped in the genera Mikusa and Bocagea. To 
this type of stamen the term JMiliusiod has been applied. Intermediate between these 
two types of stamen is one where the apices of the connectives are variously modified 
into hammer-like or sausage-like processes. Examples of this are to be found in the 
genera Clathrospermum and Popowia. To some extent (as in the Uvariew) characters taken 
from the structure of the stamen are found to be constantly co-related to certain 
characters in other organs. In other cases, however (eg., in the genus Pheanthus), 
stamens with the Uvarieoid character are associated with petals like those of DMiliusa ; 
while, in the species of Orvophea, Miliusioid stamens are associated with the corollas of 
Mitrephore. In many species a certain number of the stamens are transformed into 
linear sub-petaloid staminodes. As a rule, it is the outer stamens which are thus 
modified ; but in Anazagorea and Hupomatia it is the inner which are thus altered. In 
the latter genus the anther-cells, moreover, are introrse. = 
-. Jn the altogether anomalous Eupomatia the floral disc is concave; and, on its 
concavity, the numerous pistils are arranged, the stamens being really perigynous, although 
apparently epigynous. In some of the species of Xylopi there is a slight hollowing 
of the central part of the torus; and in this pit the bases of the pistils are sunk, the 
stamens being, however, truly hypogynous. In all the other members of the family the 
floral receptacle or ortus is either convex or very slightly concave, and the gyncecium is 
most emphatically superior. With the exception of the species of onodora, where the pistils 
Ann, Roy. Bor. Garp. Catcutra Vou. IV. 
