DISCOVERY OF THE TITANOTHERES AND ORIGINAL DESCRIPTIONS 



177 



Diplacodon emarginatus Hatcher, 1895 



Cf. Protitanotherium emarginatum Hatcher, this monograph, 

 page 377 



Original reference. — Am. Naturalist, vol. 29, pp. 

 1084-1087, pi. 38, figs. 1-4, December, 1895 (Hatcher, 

 1895.1). 



Type locality and geologic horizon. — Found by J. B. 

 Hatcher "near the base of the Diplacodon elatus beds 

 [Uinta C of Osborn], in the upper Eocene or Uinta of 

 Marsh. The locality is about 8 miles north of White 

 River and 25 miles east of Ouray Agency, Utah, and is 

 locally known as Kennedy's Hole." 



Holotype. — A skull with lower jaw (Princeton Mus. 

 11242). The anterior part only of the skull is well 

 preserved. (See fig. 112.) 



Characters. — Hatcher writes: 



The present species is at once clistinguished from D. elatus by 

 its greater size, as is shown by a comparison of the length of the 

 premolar and molar series, which is* 310 millimeters in the 

 former and 242 in the latter. 



In general appearance the cranium of D. emarginatus is re- 

 markably like some of the smaller forms of Titanotherium. 



Etymology. — emarginatus, referring to the emargi- 

 nate form of the distal end of the nasals. 



Present determination. — The species is valid but 

 generically distinct from Diplacodon Marsh and is now 

 referred to ProtitanotTierium Hatcher. (See p. 377.) 



Manteoceras Hatcher, 1895 



Cf. Manteoceras Hatcher, this monograph, page 362 



Original reference. — Am. Naturalist, vol. 29, p. 1090, 

 1895 (Hatcher, 1895.1). 



Type species. — By designation Telmatotherium valli- 

 dens (of Osborn, not Palaeosyops vallidens Cope) = 

 Manteoceras manteoceras Hay ex Osborn, MS., 1902. 



Hotelier's description — Hatcher writes: 



The genus Telmatotherium as it now stands should be divided, 

 since it embraces at least three distinct forms. The type of T. 

 vallidens should be removed from that genus and made the 

 type of a new genus. This new genus may be called Man- 

 teoceras, as suggested by Wortman from the field; it would be 

 distinguished from Telmatotherium by the absence of the infra- 

 orbital shelf, the stronger and more expanded zygomata, and 

 the concave superior aspect of the skull and incipient fronto- 

 nasal horns. 



In the above passage the reference to "the type of 

 T. vallidens" if taken by itself would lead one to regard 

 "Palaeosyops" vallidens Cope as the type of the genus 

 Manteoceras Hatcher. But a careful study of 

 Hatcher's full text and a knowledge of the history 

 of the subject proves that Hatcher had in mind the 

 "Telmatotherium vallidens" of Osborn, not of Cope: 

 because (a) Hatcher refers to his Plate 29, Figure 2, 

 as "Telmatotherium vallidens," and this figure is 

 copied from Osborn's " Telmatotherium vallidens," 

 Figure 7; (6) these figures represent Wortman's 

 original "prophet horn" skull, to which he had 

 applied the name Manteoceras "in a letter from the 



field" (Osborn); (c) the generic characters assigned by 

 Hatcher refer most clearly to this skull and are utterly 

 inapplicable to Telmatotherium {"Palaeosyops") valli- 

 dens Cope, in which only the dentition and not the 

 skull is known. 



Thus the type of the genus Manteoceras Hatcher is 

 Telmatotherium vallidens of Osborn not Cope, which 

 is equivalent to Manteoceras manteoceras Hay ex 

 Osborn MS. The generic name can not be credited 

 to Wortman, because he never published it, although 

 Osborn (1895.98), mentions it as a manuscript name. 



Etymology. — juavrtj, prophet; Ktpas, horn; in allu- 

 sion to the incipient "horns" above the orbits. 



Present determination. — This valid genus is fully de- 

 scribed on page 362. 



Dolichorhlnus Hatcher, 1895 



Cf. DoKchorhinus Hatcher, this monograph, page 396 



Original reference. — Am. Naturalist, vol. 29, p. 1090, 

 1895 (Hatcher, 1895.1). 



Type species. — Telmatotherium cornutum Osborn. 

 Characters. — Hatcher writes: 



The genus Telmatotherium as it now stands should be divided, 

 since it embraces at least three quite distinct forms * * * 

 The type of T. cornutum should also be made the type of a new 

 genus which may be called Dolichorhinus; it would be dis- 

 tinguished from Manteoceras and Telmatotherium by the 

 reduced number of inferior incisors, presence of incipient 

 horns, presence of infraorbital shelf, and position of posterior 

 nares. 



Etymology . — SoXixos, long; pis, nose. 

 Present determination. — This is a valid genus. 

 p. 396.) 



(See 



Palaeosyops ultimus Matthew, 1897 (ex Osborn MS.) 



Cf. Telmatherium ultimum Osborn, 1908, this monograph, 

 page 345 



Original reference. — Am. Naturalist, vol. 31, pp. 

 57-58, 1897 (Matthew, 1897.1). 



Subsequent reference. — Bibliography and catalogue 

 of the fossil Vertebrata of North America: U. S. 

 Geol. Survey Bull. 179, p. 631, 1902 (Hay, 1902.1). 



Doctor Matthew had no intention of describing a 

 new species. He merely stated incidentally that 

 P. ultimus, as established in manuscript by Osborn, 

 and P. paludosus both have a short-necked astragalus. 

 No type was mentioned, and the single character 

 given does not separate the species from P. paludosus. 

 Hence "Palaeosyops ultimus Matthew" (cited by 

 Hay, 1902, p. 631) remained a nomen nudum until 

 the type was fixed by Osborn in 1908. (See p. 345.) 



Etymology. — ultimus, last, latest; in allusion to the 

 relatively late geologic horizon and to the apparent 

 extinction of the race. 



Palaeosyops manteoceras Matthew, 1899 (ex Osborn MS.) 

 Cf. Manteoceras manteoceras Hay, this monograph, page 395 

 Original reference. — Am. Mus. Nat. Hist. Bull., 

 vol. 12, p. 47, 1899 (Matthew, 1899.1). 



