248 



TITANOTHERES OF ANCIENT "WYOMING, DAKOTA, AND NEBRASKA 



Group II. Retarded horn rudiments — Continued. 



3. Subfamily Palaeosyopinae — Continued. Pages 



Genus Palaeosyops Leidy (bracliycephalic, 



hyperbrachy cephalic) 157, 312, 619 



Species ffoniinalis Cope 165,317 



longirostris Earle 172,319 



paludosus Leidy 157, 319 



major Leidy 158, 321, 620 



grangeri Osborn 181, 335 



leidyi Osborn 18 1, 323, 620 



robustus (Marsh) 161, 331, 626 



copei Osborn 181, 336, 629 



4. Subfamily Telmatheriinae Osborn. "Telma- 



theres." (Middle and upper Eocene titano- 

 theres of larger size. Heads of medium length, 

 with large cingulate incisors and heavy, sub- 

 lanceolate canines; grinders large; mesatice- 

 phaUc {Telmatherium) or subbrachycephalic 

 (Sthenodectes) ; of mediportal gait.) 

 Genus Telmatherium Marsh (mesaticephalic, 



narrow sagittal crest) 160, 340 



Species cuUridens (Osborn, Scott, and 



Speir) 168, 341 



validum Marsh 160, 344 



altidens Osborn 184, 351 



ultimum Osborn 184, 345 



Genus Sthenodectes Gregory (mesaticephalic 



to subbrachycephalic) 190, 353 



Species incisivus (Douglass) 185, 354 



Group III. Accelerated horn rudiments : 



5. Subfamily Manteoceratinae (Brontopinae) Osborn. 



" Manteoceratines " (prophet-horn), "bronto- 

 pines." (Precociously horned titanotheres, of 

 the same stock as the Dolichorhininae. Skull 

 mesaticephalic, face abbreviate; feet abbre- 

 viate, brachypodal, gait graviportal; premolars 

 retarded, incisors rounded; ancestral or related 

 to the Brontops phylum of the Oligocene.) 

 Genus Manteoceras Hatcher (horns rudi- 

 mentary) 177, 362, 631 



Species manteoceras Hay 177, 365, 631 



washakiensis Osborn 182,371 



uintensis Douglass 186, 372 



Genus Protitanotherium Hatcher (horns elon- 

 gate, oval, more prominent than in Man- 

 teoceras) 176, 375 



Species emarginatum Hatcher 177, 377 



superhum Osborn 185, 379 



Genus Brachydiastematherium Bockh and 

 Maty (large size; upper Eocene of Tran- 

 sylvania) 166, 382 



Species transilvanicum Bockh and 



Maty 166, 382 



6. Subfamily Dolichorhininae Riggs. "DoUcho- 

 rhines" (long-snouted). (Middle and upper 

 Eocene titanotheres. Typically dolichocepha- 

 lic and dolichopic; nasals typically long and 

 expanding distally; precocious horn rudiments; 

 infraorbital shelf usually conspicuous.) 

 Genus Eometarhinus Osborn (ancestral to 



Metarhinus, primitive, mesaticephalic). 200, 419 

 Species huerfanensis Osborn 200, 420 



Group III. Accelerated horn rudiments — Continued. 



6. Subfamily Dolichorhininae — Continued. Pages 



Genus Mesatirhinus Osborn (ancestral to 



Dolichorhinus; subdolichocephalic) . 182,387,636 



Species Junius (Leidy) 159, 388 



megarhinus (Earle) 170, 388 



peiersoni Osborn 182, 389, 641 



Genus Dolichorhinus Hatcher (extremely 

 dolichocephalic, cyptocephalic; becoming 



extinct) 177, 396, 645 



Species superior (Riggs) 190, 395, 405 



longiceps Douglass 188,406,651 



vallidens (Cope) 162, 401 



heterodon Douglass 187,416 



intermedins Osborn 1 84, 405 



hyognathus (Osborn) _ 169, 173, 409, 646 



Jluminalis Riggs 191, 417 



Genus Metarhinus Osborn (dwarfed, aber- 

 rant, mesaticephalic) 183, 420, 648 



Species earlei Osborn 183, 420 



fluviatilis Osborn 183, 421 



cristatus Riggs 191, 429 



riparius Riggs 191, 429 



Genus Sphenocoelus Osborn (little known; (?) 



branch of Mesatirhinus) 174, 417 



Species uintensis Osborn 175,419 



7. Subfamily Rhadinorhininae ( = ?Megaceropinae) 



Osborn. " Rhadinorhines " (slender - nosed) . 

 (Middle Eocene titanotheres. Mesaticephalic, 

 cyptocephalic; infraorbital shelf reduced.) 

 Genus Rhadinorhinus Riggs (nasals short, 

 pointed; possibly ancestral to the Oligo- 

 cene Megacerops; cyptocephalic) 192, 430 



Species abbotti Riggs 192, 430 



diploconus (Osborn) 173, 431 



Group IV. Short-horned: 



8. Subfamily Diplacodontinae (=?Menodontinae, 



Brontotheriinae) Osborn. " Diplacodonts. " 

 (Upper Eocene ancestors of the Oligocene sub- 

 family Menodontinae. Heads probably mesa- 

 ticephalic; grinding teeth foreshadowing the 

 menodont type.) 

 Genus Diplacodon Marsh (horns well devel- 

 oped) 166,439 



Species eZa^Ms Marsh 166,439 



Genus Eoiitanotherium Peterson (horns well 



developed) 196, 435, 656 



Species osborni Peterson 195, 435, 656 



in. TITANOTHERES OF LOWER OIIGOCENE TIME 



(Face extremely abbreviate) 



Group I. Short-horned: 



9. Subfamily Brontopinae (Manteoceratinae) Osborn. 



" Brontopines. " (Lower Oligocene and possibly 

 middle to upper Eocene titanotheres. Progres- 

 sively brachycephalio, with short-crowned teeth 

 and moderately short feet; horns short, sub- 

 oval; incisor teeth persistent, rounded crowns, 

 one or two pair; premolars with retarded 

 tetartocones.) Pages 



Genus Teleodus Marsh (with three lower 



incisors; basal Oligocene) 227,481 



Species avus Marsh 228, 481 



primitivus (Lambe) 235, 482 



