44 CALIFORNIA ACADEMY OF SCIENCES. 
connecting the yarious organs with the body-wall and require here no particular men- 
tion. 
In the other clitellar somites we find bands of transverse muscles which 
stretch diagonally (fig. 118) across the body wall from the ventral part, almost im- 
mediately below the ventral ganglion, to the vicinity of the outer sete. The ends of 
these muscles penetrate between the lobes of the longitudinal muscular layer (fig. 
118, 7. m.) These muscles are not continuous through the somite but are grouped in 
bands. 
Ventral papille (figs. 120, 125 to 131). Argilophilus is readily distinguished 
from other California Eudrilide so far known, by its ventral papillee, occupying the 
ventral side of the intersegmental grooves from the spermatheca to the segments next 
posterior tothe clitellum. In Argilophilus marmoratus ornatus we meet sometimes with 
as many as 7 or 8 pairs, while in A. m. papillifer we find as high a number in a single 
median row, under the ventral ganglion. In no instance did I find a papilla on the 
segment itself, all invariably occurred rising from the groove between the somites, 
being in other words intersegmental. From cross-sections it will be seen that the 
papilla consists of two distinct parts, one exterior and lateral, consisting of elongated 
hypodermic supporting cells, which more or less fully enclose the interior main body 
of the papilla. This central part consists of larger or smaller gland-like bodies, vary- 
ing in size and number according to the size and age of the papilla, ete. In some 
papille these bodies fill the larger part of the papilla, in others they are confined to 
the bottom or very center of the organ. As to the nature of these organs I am, as yet, 
somewhat in doubt, though they certainly must be considered as sensory organs 
of some kind. Stained with osmie acid the bodies present in sections a darker center, 
which appears of ganglionic nature, around which are grooped larger sacs which 
again are composed of smaller, light-refractive granules, giving the idea of a reticu- 
lated protoplasm. Numerous nerve fibres connect with those bodies, and evidently 
penetrate to the gaglionic center. Long tube-like cells butt directly on these gland- 
ulous bodies, while others grooped in bundles penetrate between them connecting with 
nerve ganglia. The transition between the supporting cells and the drainpipe cells 
of the papilla is sudden and not gradual. In fact the line of demarcation between the 
two is quite prominent. The supporting cells in the young papilla, entirely enclose 
the papilla, while in the larger and full grown papilla, they are pushed towards the 
side leaving the whole center to the drainpipe cells and the glands. 
When stained with safranine the nuclei of the glandular cells become prominent, 
less so when stained with hematoxylon. The nuclei vary greatly in size and number. 
In some of the glandular bodies there are from one to three nuclei,in others none. In 
sections there are always less nuclei than divisions (cells?) apparently lying on the top of 
the reticulated mass. The nuclei may be seen directly above the nerve center, but 
generally they are found outside of it. It seems as if these glandulous bodies are 
modified agglomerations of hypodermic glands. 
The arrangement of the glandulous bodies and the tube-like cells is very 
