80 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. I28 



GIROUDIA Vercammen-Grandjean, 1952 



1952. Giroudia Vercammen-Grandjean, Ann. Soc. Beige Med. Trop., vol. 33, 

 No. 6, pp. 643-644. (Generic description in detail, classification, ex- 

 planation of name and designation of genotype: Giroudia longiscutul- 

 lata Jadin and Vercammen-Grandjean, 1952.) 



1952. Giroudia Jadin and Vercammen-Grandjean, Ann. Soc. Beige Med. 

 Trop., vol. 32, No. 6, p. 637. (Brief generic description, explanation of 

 name and designation and description of genotype: Giroudia longiscu- 

 tullata Jadin and Vercammen-Grandjean, 1952.) 



The genus Giroudia includes two unusual gahrliepiines from the 

 Belgian Congo. The above citation of Vercammen-Grandjean alone 

 as the author of the genus is based upon the obvious intention of both 

 Jadin and Vercammen-Grandjean rather than upon a strict interpreta- 

 tion of the rule of page priority which would result only in confusion. 

 Our citation is in accordance with paragraph 123, Article 28, of the 

 Copenhagen Decisions on Zoological Nomenclature (1953), under 

 which a decision to ignore page priority in special cases may be made 

 by the first reviser. 



The presence of well-developed dorsal teeth on the chelicerae serves 

 to separate Giroudia from Gahrliepia, although the other characters 

 listed by Vercammen-Grandjean (distance between usurped scuta! 

 setae, presence of well-developed eyes, and large size of scutum) 

 apply equally well to certain of the species considered herein by us 

 as Gahrliepia {Gahrliepia). 



It is pertinent to discuss some of Jadin and Vercammen-Grand- 

 jean's concepts regarding chaetotaxy. Owing to the limited number 

 of species of Gahrliepia available to these authors for study, they 

 designated as "posterolateral setae" those setae which we call PPL-i, 

 while the more anterior pair, which we call posterolaterals, were 

 designated as "mediolaterals." In their opinion, the determining fac- 

 tor regarding the homology of the PL setae is the level of the maxi- 

 mum breadth of the scutum. The extensive series of Gahrliepia in 

 our possession includes setal patterns never seen by Jadin and Ver- 

 cammen-Grandjean, and supports our contention that the second pair 

 of scutal setae are the true posterolaterals, regardless of position. 

 Thus, in G. exilis, G. laciniata, and certain other species, there is no 

 pair of setae at the level of the maximum breadth of the scutum, 

 while the PLs are in the site of the "mediolateral setae" (figs. 5 and 

 no). V/hat we call PLs are in this same position in many species 

 which also possess usurped setae at the point of maximum breadth. 

 The second pair of scutal setae, regardless of position, almost invari- 

 ably are definitely longer and stouter than the usurped setae, and this 

 is another argument for our point of view. In G. gemina, new species, 



