86 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. I28 



they are true denizens of the uncut jungle because these forms were 

 never taken in secondary vegetation or scrub terrain despite particu- 

 larly intensive collecting therein by the U. S. A. Typhus Commission, 

 the U. S. Army Medical Research Units, and the Colonial Office 

 Medical Research Unit, The following are therefore considered in- 

 habitants of primary forest : G. exilis, new species, G. netcrella, new 

 species, G. gemina, new species, G. evansi, new species, G. picta, new 

 species, G. laciniata, new species, G. tessellata, new species, G. crenu- 

 lata, new species, G. cetrata Gater, 1932, G. ornafa Womersley, 1952, 

 G. decora Womersley, 1952, G. insigne Womersley, 1952, and G. rutila 

 Gater, 1932. The following chiggers are known only from the moun- 

 tain forests of Borneo or Malaya (elevation approximately 4,500- 

 5,500 feet) : G. laciniata, tessellata, am^pullata, penetrans, granulata, 

 and cetrata. Ten of the eleven scrobiculate ("ornate") species (all 

 except G. tuberculata, new species) are included in this list of jungle 

 forms. 



SUMMARY 



The subgenus Gahrliepia includes 33 species, of which 3 are African 

 and the remainder from Asia. Eighteen new species, from Burma, 

 Assam, Thailand, Malaya, and Borneo are described and figured 

 in detail. All previously known species are discussed, and critical 

 structures are illustrated where possible. A key to the species is 

 included, Gatcria Ewing, 1938, is considered an outright synonym 

 of the subgenus Gahrliepia as here defined. It is pointed out that the 

 rarity of many of the species of Gahrliepia may be apparent rather 

 than real, owing to our ignorance of the true hosts and particular 

 sites of attachment of the chiggers. Thus, one new scrobiculate 

 ("ornate") chigger, G. (G.) laciniata, was found in abundance only 

 after it was noted that, unlike most trombiculids, this chigger did 

 not attach to the ears, but instead attached at the bases of the stout 

 spiny hairs on the muzzle of the host, Rattus alticola. G. penetrans, 

 new species ; another scrobiculate form from Borneo, burrows com- 

 pletely into the perineum of the host, Rattus tvhiteheadi. Names and 

 data for additional new species follow, and those with a scrobiculate 

 scutum are marked with an * : exilis (primary jungles of North 

 Burma) ; fimbriata (North Burma) ; darita (Assam and North 

 Burma) ; neterella (from various jungle-inhabiting rats in Malaya) ; 

 tenella (Malaya) ; elheli (from Menetes, a ground-inhabiting squir- 

 rel in Thailand) ; tylana (from Bandicota and Menetes in Thailand) ; 

 gemina (from rats and shrews in primary jungle in North Burma) ; 

 dupliseta (Assam) ; plurisetae (from the shrew Suncus, Assam) ; 



