FISHES OF OREGON AND NORTHERN CALIFORNIA. 185 
Fin CHARACTERS AND ExTeNT OF Prickiy INVESTMENT OF 139 SPECIMENS OF CoTTUS ASPER. 
Number of specimens with— 
a ] [ee at a le 
CISI/HIBIBIEI EI ElGlalala|] 2) 2] 8] & | Extent of prickly invest- 
Locality. a/e Beemer perso ee | aes Soll WEP bo seb] I ment of skin. 
7) = = — =" — = s S 
eile esueenipeive: |e | Ss ole) a iouecaltenlc 
2IZ15/8/s\s/5\/S/8|/8)a!a/8/3/3/8 
Cl/Slislsisgijal/sala}/aj a] al] a] al] a] aw] & 
PIVliaoltmalalel[aul|aleolnlJaoljalwil|ol|nio 
Dlala lA lA IAIN AlAlAlAlAlAlAlAaAlaA 
| 
Lake Washington, ]} 2) 15}. 2) 8) 7%]. 4)12) 1 2) 5} Entire upper parts except 
Wash. | caudal peduncle. 
Columbia River,Astoria.| 1) 5] 1 tty op RT 4) 3|. 5 | 2 Do. 
The Lakes, Oregon City-| 1) 9 |. mel Pate at 5) 6 35 peel Do. 
Nestucca River........-- Reseed te UP Se| soc] jh 4 Bel er: Wiis Pai elie (aol SS 9 |.........| Entire upper parts except 
| | | | caudal peduncle; area of- 
| | | | ten reduced to size of pec- 
ee be) ea toral fin. 
Takenitch River ......-. eee) wiki Bem Bo )) SI ali) | eas eat) ee ee) ae ell) Gee Do. 
Coquille River. | bates rely Goh |e) PSimetes Vay dal Fa Lah cay Wer HF CA Do. 
Flores Creek... 12 DO Gy libs 6| 5] 1|....! 6] 3] 3] Area size of pectoral or 
| somewhat larger. 
Rogue River, near |...) 8 |-.--|----j.-.. Pe ta a Tg ea 6 | Uy cee! ae Oe Soya b Entire upper parts, except 
mouth. | | caudal peduncle, often re- 
| | | duced to area no larger 
| than pectoral, 
Smith River............ oT are | 3]----| 2) 2]. Do. 
Mad River. .-... eal ee US| erat hy wtlaliec a4 Do. 
Redwood Creek 1} 3}. VA Bee See tae |e Area size of pectoral. 
Eel River. ...-- 2) Sea lee a al eee | ae Do. 
Garcia River.....-.....- Ja2=-| Gf. Pd Real | eile Entire upper parts except 
| caudal peduncle. 
Navarro River. .-....-.-- 28 aits 2 6] 5 Do. 
GualalaRiver........... Peak ets 3 | 4) 5 Do. 
PaperMillCreek,Marine | 1] 8 9} 1 Entire upper parts, except 
ounty, Cal. | | caudal peduncle, often re- 
| | duced to area half the size 
| of pectoral. 
| t 
36. Cottus gulosus (Girard). 
Cottus gulosus has been confused by recent authors with Cottus asper Richardson, the name gulosus 
having been applied to examples of C. asper from the Sacramento and neighboring streams which were 
erroneously supposed to differ from representatives of the same form from the Columbia. Cottus gulosus 
or Cottopsis gulosus described by Girard @ is a species differing from C. asper, notably in having a much 
shorter anal fin. It now appears that the species C. gulosus extends northward at least to the Columbia 
and includes the form known as Cottus perplerus, specimens of which do not differ from examples of 
C. gulosus from the Sacramento. In the Klamath basin C. gulosus is represented by Cottus klama- 
thensis, which seems to be a slightly distinct form, or at least should be so considered until certain char- 
acters which now appear to be distinctive are shown to be unreliable. C. gulosus is also closely related 
to Cottus punctulatus and Cottus beldingi. A careful examination of the relationships of these forms 
will no doubt be fruitful. 
C. gulosus occurs in most of the coastwise streams between the Sacramento and Columbia rivers, 
with possibly the single exception of the Klamath. It is usually to be found in the upper courses of the 
rivers, although it is not uncommonly associated with C. asper and C. aleuticus farther downstream. 
In @. gulosus the preopercle is always armed with one strong spine which is curved or pointed 
upward, the size of the spine, its curvature, and the angle of inclination varying somewhat in different 
individuals. It is often nearly cylindrical in shape, while again it may be rather broad and flat, 
possibly inclining toward the cylindrical form in southern examples, and being more often flat in the 
northern ones. There is also present a second comparatively weak spine which varies considerably 
in size, being sometimes reduced to a mere prominence or in rare instances disappearing entirely. 
Occasionally a third spine appears below the others. The palatine bones are usually without teeth, 
although in some cases a small patch of minute teeth may be seen. The presence or absence of teeth 
a Cottopsis gulosus Girard, Proceedings Academy Natural Sciences, Philadelphia, vol. vu, 1854, p. 129. 
